Clinobiantes paradoxus Roewer, 1927

Figs 1, 3–6; Table 1

Clinobiantes paradoxus Roewer, 1927: 302, fig. 17.

Clinobiantes paradoxus – Staręga 1992: 323.

Type material

Lectotype here designated CAMEROON • ♂; Bibundi; SMF 9900064 (examined) (Fig. 2).

Paralectotype CAMEROON • ♀; same data as for lectotype; SMF 9900064 (examined) (Fig. 2) .

Remarks

A second female is morphologically different and not conspecific with the male lectotype of Clinobiantes paradoxus or the female paralectotype herein designated (Fig. 2).

Redescription

Male (lectotype, SMF 9900064)

BODY MEASUREMENTS. Total body length 2.48, carapace length 0.76, scutum magnum length 2.17, carapace maximum width 1.36, abdominal scutum maximum width 1.8. Appendage measurements in Table 1.

DORSUM. Outline slightly hourglass-shaped (almost rectangular) with Eta (η) shape, with a very slight constriction at sulcus I level (Figs 1A, 3A). Carapace granulated, wider than long, anterior border slightly convex and unarmed (Figs 1A, 3A). Cheliceral sockets not marked (Figs 1A, 3A). Eyes separated near sulcus I (Figs 1A, 3A); interocular area granulated (Figs 1E, 3A). Carapace in lateral view straight at anterior region and slightly higher posteriorly (Fig. 1E). Abdominal scutum in lateral view convex (Fig. 1E). Sulcus I deep and well-marked, in dorsal view curved to the anterior body region (Figs 1A, E, 3A). Mesotergal areas coarsely granulated and well-defined, with sulci II–V marked but shallower than sulcus I; medially sulci II–III slightly curved to the anterior body region; sulci IV–V straight (Figs 1A, E, 3A). Mesotergal areas III–IV medially with two conical and pointed tubercles, strongly inclined backward (Figs 1A, E–F, 3A). Mesotergal area V granulated, medially with three conical and pointed tubercles. Lateral borders of abdominal scutum with rows of granules (Figs 1A, E, 3A). Free tergites granulated; free tergite I medially with three small conical tubercles (Figs 1A, E, 3A).

VENTER. Coxa I with some setiferous tubercles in anterior margin (Fig. 1C); coxa II incrassated, of same size as (or slightly larger than) coxa IV (Fig. 1B–D); anteroposterior borders of coxa III with a row of strong granules connecting with coxae II and IV, respectively (Fig. 1B–D). Posterior border of spiracular area and free sternites I–V with a row of granules; anal operculum with few granules (Fig. 1D, F). Spiracles not concealed (Fig. 1B, D).

CHELICERA. Basichelicerite unarmed, with an elongated and slightly marked bulla (Figs 1E, 3D). Cheliceral hand with sparse setae (Fig. 3D–E). Fixed and movable fingers with small triangular-shaped teeth (Fig. 3E).

PEDIPALP. Coxa elongated (i.e., remarkably longer than trochanter), proximally with one dorsoectal and one ectoventral granule (Figs 1A, C, 3A). Femur straight, proximally with a slight ventral narrowing followed by a small ventral spine (Figs 1E, 3B–C). Patella elongated, club-shaped, with a distal ventromesal spine (Figs 1E, 3B–C). Tibia with two ventromesal and two ventroectal long spines (Fig. 3B–C). Tarsus with two ventromesal and two ventroectal long spines (Figs 1E, 3B–C).

LEGS. Femur II fusiform dorsally swollen (Figs 1A, B, 6B). Tibia II slender, slightly distally swollen, with ventral small triangular tubercles (Fig. 6D). Metatarsus II with a ventrally swollen astragalus bearing two rows of triangular tubercles; astragalus-calcaneus boundary marked by a strong constriction, and slightly thickened calcaneus giving a peculiar form to the distal part of the metatarsus (Fig. 6F). Tarsi III–IV with a dense scopula. Tarsal formula: 3(2):5(3):5:5.

COLOR (in 80% ethanol). Body and appendages uniformly yellowish-brown (Figs 1A–F, 6B, D, F).

MALE GENITALIA. Penis with clearly defined boundaries between pars basalis and pars distalis (Fig. 4A). Pars basalis tubular, basally thin, broadens medially, with distal constriction (Fig. 4A). Pars distalis swollen, with maximum width at titillator level (Fig. 4A–B, D). Apical edge laminar (i.e., dorsoventrally flat) with a medial U-shaped cleft dividing it into two rounded halves (Fig. 4B, D); halves less chitinous apically, and could possibly be inflated by hemolymph pressure (Fig. 4C–D). Pars distalis with a distal depression in the ventromedial region (Fig. 4D). Each side of pars distalis armed with short, conical microsetae arranged irregularly, extending from the dorsolateral to the ventrodistal region (Fig. 4B–D). Capsula externa with two broad titillators covering most capsula interna (Fig. 4B–C). Capsula interna with two complex conductors and one stylus, basally fused. Each conductor with two medial laminar folds apically, one short dorsal and one ventral longer, visible ventrally within the U-shaped cleft; each conductor also with one broad lateral projection (Fig. 4B); stylus tubular, with its free tip fully covered by conductors (Fig. 4B–D).

II 0.525.171.253.844.022.2417.04♀ SMFLeg III0.392.470.671.693.081.079.37Leg IV0.503.470.872.204.391.2112.64

Female (paralectotype, SMF 9900064)

BODY MEASUREMENTS. Total body length 2.82, carapace length 0.76, scutum magnum length 2.31, maximum carapace width 1.38, abdominal scutum maximum width 1.88. Appendage measurements in Table 1.

BODY. Female resembles male in armature of scutum magnum, but with tubercles of mesotergal areas III–IV, posterior border, and free tergite I slightly smaller (Fig. 5A–B vs Fig. 1E). Leg II with femur, tibia, and metatarsus neither swollen nor armed (Fig. 6A, C, E vs Fig. 6B, D, F); metatarsus II thin and unarmed, with pseudoarticular rings (Fig. 6E vs Fig. 6F). Tarsal formula 3(2):5(3):5:5.

FEMALE GENITALIA. Ovipositor cylindrical (Fig. 5C), distally bearing two lobes (furca) (Fig. 5C–D, F). Each furcal lobe with five long, pointed setae (Fig. 5E) – three dorsal and two ventral – resulting in a total of six setae on the dorsal region (Fig. 5D) and four on the ventral region (Fig. 5F). External surface of furcal lobes with several short, pointed projections, irregularly distributed (Fig. 5E). Receptacle chambers located near the base of the furcal groove (Fig. 5D–F).

Distribution

Known only from the type locality (Fig. 40).