Problema Skinner & R. Williams, 1924 is a subgenus of Atrytone Scudder, 1872

Currently monotypic genus Atrytone Scudder, 1872 (type species Hesperia iowa Scudder, 1868) is sister to Problema Skinner & R. Williams, 1924 (type genus Pamphila byssus W. H. Edwards, 1880), a genus of two species, including also Hesperia bulenta Boisduval & Le Conte, [1837]. All three species are close in their genomic sequences (Fig. 15). Their genetic divergence is more consistent with them being congeners. Therefore, we treat Problema as a subgenus of Atrytone to form new or revised combinations Atrytone byssus and Atrytone bulenta.