Agraulis Boisduval & Le Conte, [1835] is a subgenus of Dione Hübner, [1819]

Monotypic genus Agraulis Boisduval & Le Conte, [1835] (type species Papilio vanillae Linnaeus, 1758) is a close sister to Dione Hübner, [1819] (type species Papilio juno Cramer, 1779) (Fig. 7). COI barcode difference between A. vanillae and Dione moneta Hübner, [1825] is 7.9%. Timecalibrated nuclear genomic tree shows that genetic divergence (Fig. 7) between Agraulis and Dione is nearly the same as the divergence among species of Boloria Moore, 1900 (type species Papilio pales [Denis & Schiffermüller], 1775) and smaller than the divergence between Heliconius Kluk, 1780 (type species Papilio charithonia Linnaeus, 1767) and Eueides Hübner, 1816 (type species Nereis dianasa Hübner, [1806]) (Fig. 7). Therefore, we treat Agraulis as a subgenus of Dione.