Ceratophyus sinicus Zunino, 1973 [中华角粪金龟]

Ceratophyus sinicus Zunino, 1973: 17, figs 4, 5, 6B (original description); Medvedev & Nikolajev 1974: 121 (distribution, key); Hillert & Král 2013: 183 (distribution).

Ceratophyus polyceros: Ren et al. 2019: 155 (distribution).

Type locality. “ Pékin [Beijing]” .

Published records. Medvedev & Nikolajev (1974): “ Peking [= Beijing]” (Beijing); “Alashan desert, Amdo and Kanssu [= Gansu]”, all with no locality specified; Hillert & Král (2013): Ertala, Haibu env., Wulan [= approx. Xireg] (Qinghai); Shanxi and Xinjiang, both with no locality specified; Ren et al. (2019): Mahuang, Pengyang Shan, Taole and Yanchi (Ningxia) (all as Ceratophyus polyceros (Pallas, 1771) . See gazetteer (Table 1) and map (Fig. 19).

Material examined (76 specimens). China, Gansu: 1 ♂ (NMPC), Heshui, 1.v.1996; Hebei: 2 ♀♀ (IZAS), Chongli, Changdi, 23.v.1981, Shibo Tian lgt.; 1 ♀ (HBUM), Weichang Xian, Chazi Linchang, 7.viii.1998, Zhe Li lgt., HBU(E)082401; 1 ♂ (HBUM), Weichang, Yudaokou Xiang, 13.-18.vi.2004, Chao Gao lgt., HBU(E)082577; 1 ♀ (HBUM), Guyuan Xian, Laozhanggou Linchang, 24.-26.vii.1998, Ming Bai lgt. HBU(E)082578; Inner Mongolia: 1 ♂ (IZAS), Xilinhot, Shuiku, 27.vii.1971; 1 ♂ (IZAS), Xilinhot, 29.vii.1971; 1 ♀ (IZAS), Juu Uda Meng [= Chifeng], Hexigten Qi, 29.viii.1974; 1 ♂ (IZAS), Alxa Meng, Alxa Zuoqi, 12.vi.1986, Q11698; 1 ♂ 1 ♀ (IZAS), Ulanqab Meng, Qahar Youyi Qianqi, 28.viii.1986, Jie Xing lgt., Q11694 [♂], Q11697[♀] ; 1 ♀ (IZAS), Jirem Meng [= Tongliao], Holingola, 6.vii.1987, Shan Bao lgt., Q42893; 1 ♂ (IZAS), Ih Ju Meng, Qianqi [= Ordos, Otog Qianqi], 7.viii.1987, Jun Kang lgt., Q42894; 1 ♂ (IZAS), Xilin Gol Meng, Xi Ujimqin Qi, 11.viii.1987, Jiang Mei lgt., Q42884; 1 ♀ (IZAS), Jirem Meng, Hure Qi, 20.viii.1987, Bingwen Gong lgt., Q42892; 1 ♂ (IZAS), Hexigten Qi, Jingpeng, 10.vii.1990, Youwei Zhang lgt.; 3 ♀♀ (HBUM), Duolun Xian, 29.vi.1994, Ren lgt. HBU(E)082581-583; 1 ♀ (IZAS), Alxa Meng, Alxa Zuoqi, Toudaoshazi, Host plant: Hedysarum scoparium, 13.viii.2011 , Xin Zhao lgt. 1 ♂ 2 ♀♀ (HBUM), Ordos, Otog Qianqi, Sanduandi Xiang, 31.v.2013, Yu Zhao & Xinglong Bai lgt.; 1 ♂, 1 ♀ (CAUC), Ih Ju Meng [= Ordos], 1972, CAUCF000002, CAUCF000003; 1♀ (CAUC), Inner Mongolia, 1978, Jia Ren lgt., CAUCF000001; Ningxia: 1 ♂ 1 ♀ (HBUM), Yanchi Xian, vi.1986, Ren lgt. HBU(E)082584 [♀], HBU(E)082585 [♂] ; 1 ♀ (HBUM), Yanchi Xian, Mahuashan, viii.1986, Ren lgt. HBU(E)082586; 3 ♂♂, 2 ♀♀ (HBUM), Taole Xian, 24.viii.1987; Qinghai: 1 ♂ (NMPC), Ertala env., 3800 m, 22.-23.vii.1992, J. Kaláb lgt.; 1 ♂, 1 ♀ (NMPC), Wulan, 3500–4000 m, 1.–6.vii.1995, A. Wrzecionko lgt.; 1 ♂, 1 ♀ (NMPC), 7 km NE of Ulan [= ap- prox. Xireg], 3020 m, 36°57.6’N 098°30.6’E, 7.vii. 2005 , individually in sheep excr., sand dune [Ch 13], J. Hájek, D. Král & J. Růžička lgt.; 13 ♂♂, 13 ♀♀ (NMPC), Haibu env., 3190–3270 m, 36°48.4-49.8’N 100°45.4-49.7’E, 13.–15.vii., 2005, individually under excr. of Procapra przewalskii, sand dune close to the bank of the Qinghai Hu lake [Ch 18], J. Hájek, D. Král & J. Růžička lgt .; Shaanxi: 1 ♂, 1 ♀ (IZAS), Yulin, 1978, Oenyao Zhu lgt .; Shanxi: 1 ♂ (NMPC), “Shansi [= Shanxi], Showchow [= Shuozhou]”, iv.1934; 1 ♂ (IZAS), Tianzhenm, 1979; 1 ♂ (IZAS), Yanggao, 13.v.1985; Xinjiang: 1 ♂ (NMPC), “Nordwestl. China, Chingkiang [= Xinjiang], Col. Reitter ”. Not located: 1 ♂ (IZAS), “Sinhouotchieng | Ordos S.E., 8.vii.1932, Licent lgt.” .

Additional material examined. Mongolia, Dornogovi: 1 ♂, 2 ♀♀ (NMPC), 1 ♂ (JSCP), Zaamyn Uud—rail- way station, 5.viii.1990, at light, D. Král & V. Kubáň lgt .

Distribution. China (Beijing, Gansu, Hebei, Inner Mongolia, Ningxia, Shaanxi, Shanxi, Qinghai and Xinjiang), see gazetteer (Table 1) and map (Fig. 19) and Mongolia (Dornogovi) (Zunino 1973; Medvedev & Nikolajev 1974; Hillert & Král 2013; Ren et al. 2019). First province records from Hebei, Ningxia and Shaanxi (China).

Note. The species Ceratophyus polyceros is recorded in “The fauna of the beetles from Ningxia, China ” (Ren et al. 2019). From the material recorded in this monograph we have examined the following specimens deposited in HBUM: two specimens from Yanchi (Ningxia), one specimen from Mahuang Shan, Yanchi (Ningxia), and five specimens from Taole (Ningxia). These specimens clearly belong to the species C. sinicus . Only one specimen from Pengyang (Ningxia) has not been found, but from the zoogeographical point of view it should be C. sinicus also.