Chamalycaeus crispatus (Godwin-Austen, 1871)

Fig. 7

Alycaeus crispatus Godwin-Austen, 1871: 91–92, pl. 4 fig. 1.

Alycaeus crispatus var. rywukensis Godwin-Austen, 1914: 373–374, pl. 154 figs 3, 3a. Syn. nov.

Alycaeus crispatus var. minimus Godwin-Austen, 1914: 373, pl. 148 figs 5, 5a. Syn. nov.

Alycaeus crispatus – Godwin-Austen 1875: 8, pl. 4 fig. 3; 1914: 371–372, 389, pl. 145 figs 1, 1a–b. Alycaeus (Dicharax) crispatus – Kobelt 1902: 367. — Gude 1921: 242–243.

Alycaeus (Dicharax) crispatus var. minima – Gude 1921: 243–244.

Alycaeus (Dicharax) crispatus var. rywukensis – Gude 1921: 244.

Chamalycaeus (Dicharax) crispatus – Ramakrishna et al. 2010: 58.

Dicharax (?) crispatus – Páll-Gergely et al. 2020: 82.

Dicharax (?) crispatus minimus – Páll-Gergely et al. 2020: 83.

Dicharax (?) crispatus rywukensis – Páll-Gergely et al. 2020: 83.

Dicharax crispatus – Gittenberger et al. 2024: 205, figs 24, 27.

Diagnosis

The narrow and high R3 swelling distinguishes this species from other congeners of the same species group.

Type material examined

INDIA • 5 syntypes of A. crispatus (Fig. 7A–E); Shibak, Habiang Garo Hills; NHMUK 1903.7.1.2635 • 11 syntypes of A. crispatus; same container as previous (probably same locality); NHMUK 1903.7.1.2759 • holotype of A. crispatus var. minimus (single shell mentioned in the original description: Fig. 7K–O); Habiang Garo Hills, W. Khasi; Godwin-Austen leg.; NHMUK 1906.4.4.176 • 2 syntypes of A. crispatus var. rywukensis (Fig. 7F –J); Rywuk, Garo Hills, South base of; Godwin-Austen coll.; NHMUK 1903.7.1.2637 .

Type localities

“Khasia, Jaintia and N. Cachar Hills” ( A. crispatus); “Habiang Garo Hills, West Khasi” ( A. crispatus var. minimus); “Rywuk Valley of the Garo Hills” ( A. crispatus var. rywukensis).

Differential diagnosis

Dicharax rugosus has a higher spire, and a more strongly and densely ribbed R1. See also under C. asaluensis .

Remarks

Godwin-Austen (1914: 372) stated in his previous paper (Godwin-Austen 1871: 93) that the Alycaeus crispatus variety from north Cachar was renamed A. asaluensis . The originally figured sample (Godwin-Austen 1871: pl. 4 fig. 1) is from Shibak, Gabir Valley (Godwin-Austen 1914: 372).

Alycaeus crispatus and A. crispatus var. minimus do not differ in any important shell characters, and therefore, the latter is a junior synonym of the former. Alycaeus crispatus var. rywukensis is smaller than the previous two and has a ribbed instead of glossy R3. Nevertheless, we interpret this as intraspecific variability.

Gittenberger et al. (2024) reported this species from Bhutan. We agree with their identification based on the photos they provided.

This species was included in the genus Dicharax due to the absence of spiral striation (Páll-Gergely et al. 2020). However, the shell shape and other characters agree with this species group, and therefore, it is reclassified in the genus Chamalycaeus . Furthermore, re-examination of the syntypes of A. crispatus var. makarsae revealed spiral striation on the teleoconch.