Cycloryx major (Godwin-Austen, 1893)

Figs 26, 27F–J

Alycaeus (Dioryx) granum var. major Godwin-Austen, 1893: 594 .

Alycaeus (Cycloryx) mangutensis Godwin-Austen, 1914: 377–378, pl. 146 figs 5, 5a.

Alycaeus (Cycloryx) graphicus var. variabilis Godwin-Austen, 1914: 403–404, pl. 146 fig. 4. Syn. nov.

Alycaeus (Cycloryx) thompsoni Godwin-Austen, 1914: 404, pl. 146 figs 3, 3a. Syn. nov.

Alycaeus (Cycloryx) mangutensis – Gude 1921: 282.

Alycaeus (Cycloryx) thompsoni – Gude 1921: 285.

Cycloryx mangutensis – Ramakrishna et al. 2010: 72.

Cycloryx thompsoni – Ramakrishna et al. 2010: 74.

Pincerna major – Páll-Gergely et al. 2020: 179.

Pincerna graphica variabilis – Páll-Gergely et al. 2020: 179.

Pincerna thompsoni – Páll-Gergely et al. 2020: 183.

Diagnosis

This species can be recognized by the dense R1 ribbing, the regular ribbing on R3, and the angle below the umbilicus, formed by the reflected outer peristome.

Type material examined

INDIA • 8 syntypes of A. major (Fig. 26A–E); Mangat Valley, Jiantia Hills; Godwin-Austen leg.; NHMUK 1903.7.1.2518 • 17 syntypes of A. graphicus var. variabilis (Fig. 26F–J); Lhota Naga; NHMUK 1903.7.1.2607 • 1 syntype of A. thompsoni (Fig. 26K–O); Munipur; Godwin-Austen coll.; NHMUK 1903.7.1.2550 .

Additional material examined

INDIA – Mizoram • 1 empty shell (Fig. 27F–J); Blue Mountain, Lawngtlai district; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; Ar53; NZSI LM1716 .

Type localities

“Mangut Valley, Jaintia Hills” ( A. major); “Lhota Naga ” ( A. graphicus var. variabilis); “Munipur” ( A. thompsoni).

Differential diagnosis

Cycloryx graphicus has more widely-spaced ribs than C. major, and the outer peristome does not form a sharp angle below the umbilicus in that species.

Remarks

The specimen from Blue Mountain is practically identical to the type specimens of P. major in all shell characters (size, shape, rib density). Specimens labelled as “ A. graphicus var.” from the Burroi Gorge, Dafla Hills (NHMUK 1903.07.1.2598) are also identical, although most specimens are larger.

Although slightly larger, A. thompsoni and A. graphicus var. variabilis also have similar traits. Therefore, they are treated as synonyms of Cycloryx major .