Echinops dissectus Kitag.

(Fig. 5 A-E)

Journal of Japanese Botany 11: 171 (Kitagawa 1935).

TYPE MATERIAL. — China • Liaoning, Shenyang city, Tiger Beach; 1.IX.1910; M. Nishimura s.n.; holotype; designated by Kitagawa (1933) .

PHENOLOGY. — Flowering and fruiting from July to August.

ETYMOLOGY. — This species epithet refers to the deeply lobed leaf blades.

DISTRIBUTION AND HABITAT. — China, Korea, Russia, Mongolia; the new record for Mongolia, occurs in Khuvsgul province. It grows in forest margins, moist grasslands, river margins, stony sunny slopes, 1300-1800 m (Shi & Greuter 2011).

ADDITIONAL SPECIMENS EXAMINED. — Mongolia • Khuvsgul, Northwest of Lashant Sum, intermountain sandy grassland; 15.VIII.2015; L.Q. Zhao M15-8008; HIMC[HIMC0013423] !

China • Inner Mongolia, Chifeng, Keshketeng Qi; 10.VIII.1958; Y.Q. Ma s.n.; HIMC[HIMC0028296] (photo!) .

TAXONOMIC NOTES

Echinops dissectus is morphologically similar to E. ritro L., which is known to occur in Mongolia. E. dissectus plants are taller than E. ritro, reaching up to 2 m tall, with simple stems, larger pseudocephalia (5-6 cm) in diam, and paper-thin leaves, but E. ritro stems are solitary or tufted, unbranched or apically branched, and pseudocephalia are 3.5-4.5 cm in diam, leaf mostly leathery.