Ceratocyrtis histricosus (Jørgensen, 1905) Petrushevskaya, 1971

Plate 10, Figs. 2–3.

Helotholus histricosa n. sp., Jørgensen, 1905, p. 137, pl. 16, figs. 86–88.

non Helotholus histricosa Jørgensen, Riedel, 1958, pl. 3, fig. 8.

non Helotholus histricosa Jørgensen, Benson, 1966, pl. 31, figs. 4–8.

Helotholus histricosa Jørgensen, Petrushevskaya, 1967, pl. 51, fig. 2.

Ceratocyrtis histricosa (Jørgensen), Petrushevskaya, 1971, pl. 52, figs. 2–4.

Ceratocyrtis histricosa (Jørgensen) forma A, Petrushevskaya, 1971, pl. 29, figs. 3–4; pl. 52, figs. 5–7.

Helotholus histricosa Jørgensen, Tan and Tchang, 1976, text-fig 53a–c.

Ceratocyrtis histricosus (Jørgensen) [sic], Bjørklund, 1976, pl. 8, figs. 19–24.

Ceratocyrtis histricsus (Jørgensen) [sic], Bjørklund, 1976, pl. 11, figs. 4–5.

non Ceratocyrtis sp. aff. C. histricosus (Jørgensen) [sic], Bjørklund, 1976, pl. 15, figs. 6–8.

non Helotholus histricosa Jørgensen, Casey, 1977, pl. 6, fig. 1.

non Ceratocyrtis cf. hystricosa (Jørgensen) [sic], Dumitrica, 1978, pl. 6, figs. 9 and 19.

Ceratocyrtis sp. aff. C. histricosus (Jørgensen), Dzinoridze et al., 1978, pl. 41, fig. 13.

Ceratocyrtis sp., Dzinoridze et al., 1978, pl. 41, fig. 20.

non Ceratocyrtis histricosa (Jørgensen), Takahashi and Honjo, 1981, pl. 7, figs. 6–7.

non Helotholus histricosa Jørgensen group, Benson, 1983, pl. 8, figs. 1–3.

? Ceratocyrtis histricosa (Jørgensen), Nigrini and Lombari, 1984, pl. 15, fig. 6.

Ceratocyrtis histricosus (Jørgensen), Funakawa, 1994, pl. 7, fig 1a–b.

non Helotholus histricosa Jørgensen, Chen et al., 2017, pl. 68, figs 12–13.

? Clathromitra sp. 2, Chen et al., 2017, pl. 68, fig. 4 (non figs. 7–11).

Helotholus histricosa Jørgensen, Trubovitz et al., 2020, supplementary data 7.

Remarks. Petrushevskaya (1971) transferred this species from Helotholus to Ceratocyrtis, as discussed in the genus discussion above. Since then, not all authors have recognized this name change, some of them retaining Helotholus . Many illustrated specimens of Ceratocyrtis (or Helotholus) histricosa in previous literature are typical of those observed during this study. However, there have been a number of specimens of Sethoconus tabulatus (Ehrenberg, 1873b) Haeckel, 1887, forms similar to Helotholus vema (Hays, 1965) or possibly species of Botryopera, specimens of Antarctissa and Pseudodictyophimus, as well as different Ceratocyrtis taxa that have been misidentified as Ceratocyrtis histricosus (Jørgensen), indicating multiple contradictory concepts in the literature. Here we follow the species concept and genus transfer of Petrushevskaya (1971).

Range. Late Miocene—Late Pleistocene, EEP (Table 1).