Elbenia (Elbenia)? tenera Brunner von Wattenwyl, 1878

(Figs 2B, 5–7)

Elbenia tenera Brunner von Wattenwyl, 1878: 166 — Brunner von Wattenwyl, 1891: 78; Brunner von Wattenwyl, 1898: 254; Kirby, 1906: 425; Hebard, 1922: 143.

Elbenia triangularis Brunner von Wattenwyl, 1898: 255 — Kirby, 1906: 425; Hebard, 1922: 143 (synonymisation); Tan, 2014: 333; Gorochov, 2023b: 9, 24; Tan et al., 2024a: 457.

New material examined. EAST MALAYSIA; Sabah State • 2♂; Sinipung Hill, near Long Pasia and Meligan; N4.42605 E115.72710, 1177.0± 5.3 m.a.s.l.; 20 November 2024, 13h03; resting on Melastoma leaf; coll. M.K. Tan; SBH.24.152, 153 (ZRC) • 1♂; Sinipung Hill, near Long Pasia and Meligan; N4.42801 E115.72686, 1188.0± 6.8 m.a.s.l.; 20 November 2024, 13h20; flying around in open area; coll. M.K. Tan; SBH.24.154 (FRC) • 1♂; Meligan (P28 a.k.a. pt 2); N4.57441 E115.73289, 1374.0± 6.5 m.a.s.l.; 23 November 2024, 19h49; on foliage of rattan; coll. M.K. Tan; SBH.24.234 (Fig. 5) (ZRC) • 4♂; Meligan (P28 a.k.a. pt 2); N4.57619 E115.73923, 1266.0± 6.8 m.a.s.l.; 23 November 2024, 22h00; attracted to light trap on the road; coll. M.K. Tan; SBH.24.255, 256, 259, 260 (MNHN and ZRC) • 1♀; Sinipung Hill, near Long Pasia and Meligan; N4.42267 E115.72818, 1153.0± 5.8 m.a.s.l.; 20 November 2024, 12h28; flying around in open area; coll. M.K. Tan; SBH.24.151 (FRC) .

Remarks. Our male specimens (Fig. 6) resemble the specimens described in Gorochov (2023b): the male tenth abdominal tergite extends posteriorly, forming a large hammerhead-like extension at the apex, with each end having dorsal and ventral lobes; but this extension is a little disproportional in our specimens compared to the ones by Gorochov (2023b). Their cerci are similar in being simple, small, more strongly curved subapical parts and with the very small apical parts curved medially and having the apex sclerotized and slightly falcate (Figs. 6D–F). The subgenital plate is very large, elongated, curved dorsad at the apical half, narrowly but deeply emarginated, at the apex each with two (sometimes with additional one) tooth-like denticles (Figs. 6D–F).

Type locality. Borneo.

Natural history. This species is very common at the forest edge and can be found flying around even in the day.

Calling song (Fig. 7). The calling song is an isolated echeme made up of 3–7 isolated syllables, with an average of 4±1 syllables. The average echeme duration is 0.42± 0.16 s (0.19– 0.78 s). Within each echeme, the average syllable duration is 4.0±1.0 ms (2.5–6.0 ms) and the average down time (silent interval between the two syllables) is 0.13± 0.02 s (0.09– 0.17 s). On rare occasions, doublet of syllables may occur. The frequency spectrum is broadband and is not symmetrical and has a peak frequency of 18.9±1.5 kHz (17.3–23.0 kHz).