Oedignatha tongbiguan sp. nov.

Figs 1–4

Type material. Holotype ♂ (MHBUARA-2024-882-17), CHINA: Yunnan Province: Yingjiang County, Tongbiguan Town (24.6680°N, 97.5990°E; 913.4 m a.s.l.), 5.X.2024, leg. K. Yu, Y. Ni and Y. Xu. Paratypes: 3♀ (MHBUARA-2024-882-18–20), same data as holotype .

Etymology. The specific epithet is derived from the name of the type locality, noun in apposition.

Diagnosis. Males of O. tongbiguan sp. nov. resemble O. dian Lu & Li, 2023 in having a similar shaped embolus and a distally sclerotized conductor, but can be distinguished from the latter by: 1) the retrolateral tibial apophysis bifurcated basally (Figs 2C, 4C; vs. apically bifurcated in O. dian, see Lu et al. 2023: figs 5C, 6C); and 2) embolus originating at 8-o’clock position ventrally (Figs 2B, 4B; vs. at 9-o’clock position in O. dian, see Lu et al. 2023: figs 5B, 6B). Females are most similar to the females of O. shaanxi Li & Yao, 2023 in possessing a membranous epigynal window and spherical spermathecae, but differs by: 1) the width of the epigynal window about 1.5 times its height (Figs 3A, 4D; vs. width about 3 times its height in O. shaanxi, see Chu et al. 2023a: fig. 2A); 2) copulatory ducts narrow (Figs 3B, 4E; vs. wide in O. shaanxi, see Chu et al. 2023a: fig. 2B).

Description. Male (holotype, Figs 1A–B). Total length 6.55, carapace 3.21 long, 2.09 wide; abdomen 3.34 long, 1.86 wide. Carapace oval, dark brown. Cervical and radial grooves indistinct. Clypeal hump arched (Fig. 1A). Labium, sternum and endite reddish brown (Fig. 1B). Chelicerae with three pro- and six retromarginal teeth. Eye sizes and interdistances:AME 0.13, ALE 0.14, PME 0.10, PLE 0.11; AME–AME 0.13, AME–ALE 0.17, ALE–ALE 0.70, PME–PME 0.28, PME–PLE 0.29, PLE–PLE 0.97, ALE–PLE 0.11. Legs brown. Leg measurements: Ⅰ 9.41 (2.51, 0.89, 2.35, 2.24, 1.42), II 7.97 (2.25, 0.89, 1.80, 1.89, 1.14), III 7.04 (1.99, 0.81, 1.42, 1.80, 1.02), IV 10.16 (2.69, 0.97, 2.27, 2.81, 1.42); leg formula 4123. Tibial and metatarsal ventral spines depicted in Table 1. Abdomen oval, with complete dorsal scutum (Fig. 1A).

Palp (Figs 2, 4A–C). Cymbium with narrow groove basally. Retrolateral tibial apophysis bifurcated at base, prolateral branch S-shaped in ventral view, less sclerotized than retrolateral branch, retrolateral branch triangular in retrolateral view (Figs 2C, 4C). Bulb less than half length of cymbium (Figs 2B, 4B). Embolus filamentous, originating from 8-o’clock position and extending clockwise to anterior side of conductor (Figs 2B, 4B). Conductor widely triangular, with heavily sclerotized edge (Figs 2, 4A–C).

Female (paratype, Figs 1C–D) (MHBUARA-2024-882-18). Same as male except for the following: total length 7.19, carapace 2.92 long, 2.25 wide; abdomen 4.27 long, 2.52 wide. Chelicerae with eight retromarginal teeth. Eye sizes and interdistances: AME 0.13, ALE 0.15, PME 0.10, PLE 0.11; AME–AME 0.12, AME–ALE 0.16, ALE– ALE 0.71, PME–PME 0.29, PME–PLE 0.25, PLE–PLE 0.97, ALE–PLE 0.10. Leg measurements: Ⅰ 8.91 (2.41, 0.83, 2.19, 2.14, 1.34), II 7.61 (2.09, 0.82, 1.74, 1.75, 1.21), III 6.57 (1.80, 0.79, 1.29, 1.68, 1.01), IV 9.96 (2.60, 0.85, 2.20, 2.80, 1.51). Tibial and metatarsal ventral spines depicted in Table 1.

Genitalia (Figs 3, 4D–E). Epigynal plate sclerotized, with posteriorly located membranous epigynal window (Figs 3A, 4D). Copulatory ducts short, partially visible in dorsal view. Spermathecae spherical. Bursae spherical, clearly smaller than spermathecae. Fertilization ducts pointing anterolaterally (Figs 3B, 4E).

Variation. Female (n=3): body length 6.87–7.32.

Distribution. Known only from the type locality.