Scaptomyza (Grimshawomyia)

Diagnosis. The genus Grimshawomyia was described by Hardy (1965) and contains two species, G. palata and G. perkinsi, the latter of which was initially described as a member of the genus Drosophila by Grimshaw (1901). The unique male genitalia, which feature surstyli that are exposed and an epandrium which is developed into a pair of moderately large lateral lobes, extending beyond the apices of the surstyli (Figs. 5­6), suggest that these taxa actually form a clade within the genus Scaptomyza . This clade is also characterized by having the second antennal segment sharply pointed at the apex, extending over the base of the third segment; the vertical and ocellar setae inserted into the somewhat swollen sides of the vertex; a long costal fringe, which extends nearly to vein R 4+5, and the distinctive wing markings.

Included Taxa. Scaptomyza (Grimshawomyia) palata (Hardy), comb. nov. from Maui and O‘ahu, and S. (Grimshawomyia) perkinsi (Grimshaw), comb. nov. from Maui, O‘ahu and Hawai‘i (Nishida, 2002).

KEY TO SPECIES OF THE SUBGENUS Grimshawomyia Hardy

16. Clypeus and lower margin of the face yellow; wing with distinct pattern, apex lightly infuscated; two reclinate orbital setae present on frons; coxae predominantly yellow; foretarsi brown to black; third costal section shorter, roughly 2.5 times longer than fourth; each pleuron with a broad, transverse yellow vitta, lower 1/2 of katepisternum yellow, surstyli longer than wide, each with a row of fine teeth on venter. Hawai‘i ..... ....................................................................................................... perkinsi (Grimshaw)

­ Clypeus pale brown; lower margin of the face predominantly brown, with a thin band of yellow; wing pattern similar to above, but with hyaline area at apex; three distinct reclinate orbital setae present on frons; coxae brown; foretarsi yellow; third costal section longer, about 3.5 times longer than fourth; pleura almost entirely brown, lacking distinct vittae; surstyli plainly visible, evenly rounded on ventral surface, lacking conspicuous teeth. O’ahu, Maui ................................................................... palata (Hardy)