Leopoldamys Ellerman, 1947 . Proc. Zool. Soc. Lond., 117:267.

TYPE SPECIES: Mus sabanus Thomas, 1887 .

COMMENTS: Definition and contrasts with Rattus and Niviventer provided by Musser (19816), who also reviewed morphological, chromosomal, and distributional characteristics. Leopoldamys is dentally similar to Berylmys, Maxomys, and Niviventer . Sperm morphology united Leopoldamys with Berylmys, Sundamys, and Rattus (Breed and Yong, 1986), but alliance is based on shared spermatozoal form that is likely primitive. Analyses of chromosomal traits suggested Leopoldamys is more closely related to Bandicota, Berylmys, Nesokia, Rattus, and Sundamys, than to Lenothrix, Maxomys, or Niviventer (Gadi and Sharma, 1983); biochemical data separated Leopoldamys far from Rattus (Chan et al., 1979).