Sagola waipouaensis Park and Carlton, new species

(Figs. 8b, 8f, 8i–j, 10)

Type Material. Holotype. NEW ZEALAND: Northland: 1♂ (NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: ND: Waipoua SF, Yakas Tree Tr. 350m 11–14.iv.1980 broadlf.-podocarp A. Newton, M. Thayer ”, “berl, litter at bases of Metrosideros rubusta trunks”, “ HOLOTYPE Sagola waipouaensis Park and Carlton 2013 ” . Paratypes (n = 11: 9 males; 2 females). NEW ZEALAND: Auckland: 1♂ (NZAC), Whangateau, 27 XII 1951, P625; Northland: 5♂♂ 1♀ (2♂♂, DSC; 3♂♂ 1♀, FMNH), Waipoua SF, Toronui Tr., 150m, 13 IV 1980, kauri-podocarp-broadlf.-nikau palm forest, A. Newton, M. Thayer, berl., leaf & log litter, forest floor; 1♂ (DSC), Waipoua SF, 0.9km e Forest hqtrs., 120m, 26 XI–4 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 686, leaf & log litter, forest floor; 1♂ (slide-mounted; FMNH), Waipoua SF, 0.9km e forest hqtrs., 120m, 26 XI–4 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 686, FIT & window trap ; 1♂ (DSC), Waipoua SF, 1.7km nw Waikohatu Stream Bridge, 380m, 28 XI–6 XII 1984, hdwd.-podocarp forest, A. Newton, M. Thayer 691, FIT, window trap ; 1♂ (FMNH), Waipoua SF, Kauri Ricker Track, 120m, 26 XI–4 XII 1984, kauri-podocarp-hdwd., A. Newton, M. Thayer 684, berl., leaf & log litter, forest floor; 1♀ (FMNH), Waipoua SF, kauri Ricker Tr., 100m, 11–15 IV 1980, kauri-podocarp-broadleaf, A. Newton, M. Thayer, berl., leaf & log litter, forest floor.

Etymology. This species is named after the type locality, Waipoua, Auckland.

Diagnosis. This species can be distinguished from the other species of the rugifrons speciesgroup by the following combination of characters: smaller body, length 2.1–2.6 mm; frontal rostrum with distinctive punctures; antennomere 1 approximately 2.5 times longer than wide, 4–8 longer than wide; round posterior frontal fovea without surrounding dense setae; ventral surface of male head with distinctive pair of short carinae behind eyes; shape of genitalia unique to species.

Description. Length 2.1–2.6 mm. Body brown, elytra, legs, and maxillary palpi paler (Fig. 8b). Head: As long as wide, widest across eyes (Fig. 8i). Frontal rostrum with distinctive punctures. Ventral surface of male head with distinctive pair of short carinae behind eyes (Fig. 8j). Antennomere 1 approximately 2.5 times longer than wide, 2–3 subquadrate, 4–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep and reaching midpoint of eye from frontal rostral lobes. Anterior frontal fovea round. Posterior frontal fovea round. Eye small and prominent, one-half length of temple. Female eye small one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular in male (Fig. 8b), subquadrate in female. Hind wings present in male, absent in female. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe shorter than parameres, divided in 2 lobes. Apical lobe of major lobe triangular (Fig. 8f). Minor lobe slender and forked into 3 branches (Fig. 8f). Phallobase of median lobe nearly symmetrical and rounded (Fig. 8f).

Distribution. Auckland, Northland (Fig. 10: black circles).

Habitat. Specimens of this species were collected using window and flight intercept traps, or by sifting leaf and log litter in kauri, broadleaf, hardwood, nikau palm, and podocarp forests.