Sagola rustica Broun, 1915

(Figs. 31a, 31e, 32)

Sagola rustica Broun, 1915: 285 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.

Sagola subcuneata Broun, 1921a: 488 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244. New synonymy.

Sagola brevifossa Broun, 1921a: 501 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241. New synonymy.

Type Material. Holotype. NEW ZEALAND: Mid Canterbury: 1♀ (BMNH), glued on rectangular card, “Type” [red label, printed]; “3700.♀.” [white label, handwritten]; “Rakaia. 18_5_1912.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola ♀. rustica.” [white label, handwritten]. Holotype of Sagola brevifossa: NEW ZEALAND: Otago Lakes: 1♂ (BMNH), glued on rectangular card, “4014.♂” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Routeburn. 16.2.1914.” [white label, handwritten]; “ Sagola ♂. brevifossa.” [white label, handwritten]. Holotype of Sagola subcuneata: NEW ZEALAND: Mid Canterbury: 1♂ (BMNH), glued on rectangular card, “3995.♂” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Sec x?” [white label, handwritten]; “Moa Basin . 20.10.1913.” [white label, handwritten]; “ Sagola ♂. subcuneata.” [white label, handwritten] .

Additional Material (n = 22: 5 males; 17 females). NEW ZEALAND: Fiordland: 1♂ 2♀♀, Secretary I, Gut Bay, 23 XI 1981, C.F. Butcher, sifted litter around hut 81/177; 1♂, Breaksea Sound, Gilbert I No 6, 12 III 1983, C.F. Butcher, sifted rotten wood 83/43; 1♂, South Arm, L. Manapouri, 15 XII 1998, J. Nunn, underside log with white rot; 1♀, South Arm, L. Manapouri, 13 XII 1998, J. Nunn, in moss; 1♂ 1♀, Fiordland NP, Kepler tr. below Mt. Luxmore Hut, 900m, 45°24.865′S, 167°38.675′E, 11 XII 2005, Nothofagus forest, pyt. fogging mossy log, A. Solodovnikov, D. Clarke, ANMT site 1176; 1♂, Murchison Mts, lower east McKenzie Burn, 945m, 8 XII 1983, C.A. Muir, litter in N. menziesii forest; 2♀♀, Fiordland NP, Milford Sound rd., Smithy Creek Campground area, 400m, 44°57.065′S, 168°01.156′E, 9–11 XII 2005, Nothofagus fusca & N. menziesii open forest, pyt. fogging old mossy & fungusy logs, A. Newton, ANMT site 1170; 1♀, Fiordland NP, Milford Sound rd., Gertrude Saddle car park area, 800m, 44″46.13′S, 168″00.16′E, 9 XII 2005, very mossy Nothofagus menziesii forest, FMHD #2005-087, berl., leaf & log litter, A. Newton, M. Thayer, ANMT site 1169; 1♀, Fiordland NP, Borland rd., Mt. Burns Tops tr. above Borland Saddle, 1075m, 45″44.883′S, 167″22.938′E, 10 XII 2005, subalpine tussockland, FMHD #2005-096, berl., unsifted at bases of shrubs ( Dracophyllum, possibly Fillfolium), M. Thayer, ANMT site 1173; 2♀♀, Eglinton Valley Divide, 400m, 13 II 1982, C.F. Butcher, moss/liverworts 82/29; 2♀♀, Mt. Annetta Sdle, Little Red Hill, Barrier R., 1433m, 4 II 1975, litter alpine swards 75/43; 1♀, Simonin Pass, W. Olivine Ra., 1067m, 23 I 1975, G.W. Ramsay, litter in Nothofagus forest 75/37; Westland: 4♀♀, Mt. Aspiring NP, Arawata Biv, 840m, 5 II 1989, LCNZ 89/2, J.W. Early, R.M. Emberson, scrub litter.

Diagnosis. This species can be distinguished from the other species of this species-group by the following combination of characters: larger body, length 2.6–2.8 mm; ventral surface of male head convex with setose gular region; eye large and prominent, approximately one-half length of temple; frontal sulcus deep and reverse keyholeshaped; shape of antennomeres and genitalia unique to species.

Redescription. Length 2.6–2.8 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (Fig. 31a). Head: Head round, widest across eyes (Fig. 31a). Ventral surface of male head convex with setose gular region. Male antennomere 1 approximately 2 times longer than wide, 2 subquadrate, 3–8 longer than wide, 9–10 subquadrate. Female antennomere longer than male, 2–3 subquadrate, 4–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep, reaching posterior end of eye. Anterior frontal fovea oval, posterior frontal fovea elongate. Eye large and prominent, approximately one-half length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular (Fig. 31a). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle, less developed in female. Aedeagus: Median lobe divided, minor lobe longer with tubercles (Fig. 31e). Phallobase of median lobe symmetrical and rounded (Fig. 31e). Parameres symmetrical with sparse setae from apex to midpoint, broader than median lobe (Fig. 31e).

Type Locality. Rakaia Gorge, Mid Canterbury, New Zealand .

Distribution. Fiordland, Mid Canterbury, Otago Lakes, Westland (Fig. 32: black circles).

Habitat. Most specimens of this species were collected by fogging or sifting mossy leaf or log litter in Nothofagus forests.

Comments. Specimens of S. rustica can be distinguished from other species by the shape and size of the antennomeres, frontal fovea, frontal sulcus, size of eye, and unique genitalia. The type specimens of S. brevifossa and S. subcuneata share these diagnostic characters, and these species have been collected at or near the type locality of S. rustica . For these reasons, we have placed S. brevifossa and S. subcuneata in synonymy with S. rustica .