List of Onthophagus species mentioned in this paper

Species arranged in two groups, within these groups according to approximate similarity (also in Table 1). PNG = Papua New Guinea, WNG = West New Guinea, Indonesia. Holotype depository in square brackets.

Broad-eyed: O. acerus group O. daymanus new species—PNG [AMNH] O. baiyericus new species—PNG [CMNC] O. bituberoculus new species—WNG [MBBJ] O. mimikanus Balthasar, 1969 —WNG [NMPC] Narrow-eyed: O. kokodentatus group

O. acerus Gillet, 1930 — PNG [IRSNB] O. kukali new species—PNG [CMNC] O. aceroides new species—WNG [MBBJ] O. kokodentatus new species—PNG [CMNC] O. kokocellosus new species—PNG [CMNC] O. dissidentatus new species—PNG [CMNC] O. kokopygus new species—PNG [CMNC] Ungrouped species

O. ofianus new species—PNG [CMNC]