Phanoperla simplex Zwick

Phanoperla simplex Zwick, 1982:114 .

Holotype ♂ (British Museum of Natural History). Kuala Lumpur, Malaysia

Phanoperla simplex: Zwick & Sivec, 1985:124 . In part

Material examined. Thailand: Chiang Mai Province, Chiang Dao National Park, Namtok Srisungwan, 98° 57’ N, 19° 37’ E, 600 m, 26 May 2002 , Chiang Mai University Team, 1♀ (PMSL) . Chiang Mai Province, Chiang Dao, 18 May 1983, 1♂ (PMSL) . Khampaeng Phet Province, Khlong Lan National Park, Namtok Klong Lan, 99° 16’ N, 16° 07’ E, 310 m, 19-20 October 2002, I. Sivec, 1♀ (PMSL) . Mae Hong Son Province, Namtok Mae Surin National Park, Mae Nam Pai, 97° 59’ N, 19° 21’ E, 310 m, 15 October 2002, I. Sivec, 1♀ (PMSL) . Same site, 19 March 2002, 1♂, 1♀ (PMSL) .

Remarks. Zwick (1982) reported this species from several Malaysian and Thai sites and additional localities were given in Zwick & Sivec (1985). The latter study includes specimens from Sumatra which have egg collars markedly different from mainland specimens. In our experience this much variation in such a structure suggests more than one species is involved. In our opinion various populations of P. simplex should be carefully re-examined when material becomes available.