Dacnulysia gen. nov.
Figs 1–3
Type species: Dacnulysia chaenomastax sp. nov.
Diagnosis. Head nearly square (Fig. 2 J); third antennal segment nearly as long as fourth segment, scapus apically oblique, ventrally shorter than dorsally (Fig. 2 H); eyes glabrous; clypeus semi-circular and medium-sized (Fig. 2 I); mandible considerably protruding beyond condylus ventro-basally (Fig. 2 K), first tooth right angled and forming a wide gap with middle tooth (Fig. 2 I), middle tooth acute, much longer than both lateral teeth and with an extra fourth tooth dorsally, tooth small and lobe-shaped (Figs 2 I, 2L, 3A), an oblique diagonal ridge extend from third tooth to upper basal corner of mandible (Fig. 2 I); pronope round and large; metanotum acutely protruding dorsally (Fig. 2 C); vein 3-SR of fore wing subequal to 2-SR (Fig. 2 A), vein CU1b of fore wing long, CU1a at same level as 2-CU1, pterostigma elongated and basal part subparallel-sided, vein r issued from apical quarter of pterostigma (Figs 1, 2 A); vein M+CU of hind wing much shorter than vein 1-M (Fig. 2 B); marginal cell of hind wing narrow; precoxal sulcus complete, crenulate and transverse (Fig. 2 C); lobes of mesoscutum similarly convex and smooth; surface of propodeum densely and coarsely reticulate-rugose, areola absent (Figs 2 D, 2F); metanotum acutely protruding dorsally (Fig. 2 C); dorsope absent, but position indicated by two superficial elongate depressions (Fig. 2 F); metasoma rather compressed, its second and following tergites smooth (Fig. 2 E); apex of ovipositor sheath rounded, without spine (Fig. 3 E).
Comparative diagnosis. The new genus runs in the key to world genera by Fischer (1974) to Phaenocarpa Foerster, 1863, or to Idiasta Foerster, 1863, and Hoplitalysia Ashmead 1900, because of the vein 3-SR of fore wing being nearly subequal to vein 2-SR and the metanotum acutely protruding dorsally. It differs as follows: the new genus has the dorsope absent (versus present in Phaenocarpa), head nearly square (vs distinctly transverse), middle tooth of mandible with extra protuberance (fourth tooth present, vs absent) and middle tooth much longer than lateral teeth (vs somewhat longer), and vein 3-SR of fore wing subequal to 2-SR (vs longer than vein 2-SR, rarely slightly so). The flat scutellum, the often protruding metanotum, the long oblique diagonal ridge of the mandible and the distinctly protruding ventro-basal corner of the mandible are also found in Idiasta . However, the new genus has the head nearly square in dorsal view (vs much wider than long in Idiasta), vein CU1b of fore wing long and CU1a at same level as 2-CU1 (vs CU1b medium-sized and CU1a far below level of 2-CU1), pterostigma elongated and basal part subparallel-sided (vs triangular) and vein M+CU of hind wing much shorter than vein 1-M (vs longer than vein 1-M). Differs from Hoplitalysia by its flat scutellum (vs conical in Hoplitalysia), pterostigma elongate (vs triangular), vein r emitted near apical quarter of pterostigma (vs slightly beyond middle), third antennal segment nearly as long as fourth segment (vs 0.7 ×), eyes glabrous (vs setose), notauli complete (vs absent on disc) and head nearly square in dorsal view (vs nearly twice wider than long). The nearly square head (both in dorsal and anterior view) is very rare in Alysiini and so far only known from Eudinostigma Tobias, 1986 (van Achterberg, 1988b) . Eudinostigma has head in dorsal view 1.6–2.4 × as wide as mesoscutum (vs 1.3 × in Dacnulysia), third antennal segment 1.1–1.2 × as long as fourth segment (vs slightly shorter in Dacnulysia), head in anterior view strongly transverse (vs slightly wider than high in Dacnulysia), vein r issued before middle of pterostigma (vs behind middle of pterostigma), dorsope normal (vs absent and position indicated by two superficial elongate depressions) and middle tooth somewhat longer than lateral teeth (vs much longer).
Etymology. Name is a combination of two generic names: Dacnusa Haliday and Alysia Latreille, because the new genus belongs to the Alysiini according to the venation but the shape of the body reminds of the Dacnusini. Gender: feminine.
Biology. Unknown.
Distribution. Oriental (China).