21. Leiodes okawai Nakane, 1963

Japanese name: Kiiro-ô-tamakinokomushi (Figs. 6, 64–66, 116)

Leiodes okawai Nakane, 1963: 41; DAFFNER (1983): 108 (redescription); PERREAU (2004): 197 (catalogue).

Type locality. Japan, Honshu, Mie Pref., Hirakura.

Type material examined. JAPAN: HONSHU: HOLOTYPE, ♂, Mie Pref., Hirakura, 12.vi.1960, T. Okawa leg. (HUMS).

Additional material examined. JAPAN: KYUSHU: 1♂, Kumamoto Pref., Ue Village, Mt.Shiragadake 14.viii.2000, K, Kido leg. ; 1 ♂, Kumamoto Pref., forest in Itsukimura Village (32º28′36″N 130º46′42″E, alt. 900 m), 17.vii.2009, M.Makihara leg.(PT). SHIKOKU: 1 ♂, Kôchi Pref., forest in Aki (33º39′15.2″N 134º09′28.9″E, alt. 700 m), 30.vi.2010, M. Makihara leg. (PT) ; 1 ♂, Kôchi Pref., forest in Aki (33º39′14.9″N 134º09′28.8″E, alt. 700 m), 4.viii.2010, M. Makihara leg. (MT) ; 5 ♂♂, 1 ♀, Ehime Pref., West ravine of Mt. Shiratsue, 3.vii.1986, T. Nagata leg. (collected by treacle traps) (EUMJ) ; 3 ♂♂, 1 ♀, Tokushima Pref., Yamakawa Town, Mt. Kôtsu, 2–10.viii.2003, K. Tanaka leg. (FIT) ; 2 ♂♂, 1 ♀, Tokushima Pref., Yamakawa Town, Okunoi, 3–8.vii.2003, K. Tanaka leg. (FIT) ; 1 ♂, 1 ♀, 8–14.vii.2003, same data as the former except for the date. HONSHU: 2 ♂♂, 2♀♀, Okayama Pref., Chûka Village, Mt. Yamanorisen, 11.vii.2004, Y.Fujitani leg.(FIT) ; 1♂, 3♀♀, Okayama Pref., Nagi Town, Takiyama, 29.vi.–9.vii.2005, S. Suzuki leg. (FIT) ; 1 ♂, same data as the former except for the date, 14.–23.vii.2005; 6 ♂♂, 4 ♀♀, Okayama Pref., Nagi Town, Mt. Nagisan, 29.vi.–9.vii.2005, S. Suzuki leg. (FIT) ; 1 ♂, 1 ♀, Mie Pref., Mt. Nonobori, 8.vii.2001, M. Inagaki leg. (FIT) ; 1 ♀, same data as the former except for the date, 12.vii.2001; 1 ♂, 2 ♀♀, same data as the former except for the date, 15.vii.2001; 1 ♂, same data as the former except for the date, 22.vii.2001; 1 ♂, Gifu Pref., Shirakawa Village, Ô–shirakawa, 3.vii.2004, K. Toyoshima leg. (FIT) ; 1 ♂, 1 ♀, 19.vii.2004, same data as the former except for the date; 1 ♂, 1 ♀, same data as the former except for the date, 31.vii.2004; 1 ♂, 1 ♀, Fukui Pref., Ôno City, Heikedaira, 1.viii.1997, S. Inoue leg. ; 1 ♂, Shizuoka Pref., Sakuma Town, Mt. Ryuto-zan, 7.vii.1995, T. Kishimoto leg. ; 4 ♂♂, Tokyo Pref., Hinohara Village, Mt. Mitôsan (alt. 1100 m), 16–23.vii.2008, H. Takano leg. (FIT) ; 1 ♂, 1 ♀, same data as the former except for the date, 16–23.vii.2008; 2 ♂♂, Saitama Pref., Ohtaki Village, Nakatsukawa-keikoku, Oku-Chichibu-rindô (alt. 1300m), 30.vii.–7.viii. 004, K. Arai & S. Arai leg. (FIT) ; 2 ♀♀, Saitama Pref., Naguri Village, Mt. Arimayama (alt. 1200 m), 17.–24.ix.2004, K. Arai and S. Arai leg. (FIT) ; 5 ♂♂, 1 ♀, Miyagi Pref., Sendai City, Mt. Izumigadake, Kuwanuma-rindô, 12.–22.vii.2009, M. Oikawa leg. (FIT) ; 1 ♀, Aomori Pref., Kuroishi City, Nagayasawa, 28.v.1998, T. Ichita leg. ; 1 ♂, Aomori Pref., Mutsu City, Kanauchi-chô, Yunokawa-tôge, 24.vii.1999, S. Araki leg. All specimens except for type series are preserved in FUFJ .

Diagnosis. Coloration. Dorsum usually unicolor, brown or yellowish brown; antennomeres 1–6 and 8 brown; antennomeres 7, 9, 10, and basal 3/5 of antennomere 11 dark brown; apical 2/5 of antennomere 11 light brown.

Body 2.6–3.5 mm in length, ca. 1.8× as long as wide (Fig. 64A). Head densely and minutely punctate, bearing some large punctures (Fig. 64A); antennomeres 1–3 longer than wide; antennomere 11 about as long as wide; remaining antennomeres each wider than long; antennomere 11 robust and clearly narrower than antennomere 10 (Fig. 64C). Pronotum feebly sinuate at posterior margin, densely and minutely punctate (Fig. 64A). Elytra not transversely strigose; each elytron with nine rows of punctures, bearing small number of large punctures and moderate number of very fine punctures between rows (Fig. 64D); row 9 invisible in dorsal view, almost straight, subhumeral row absent (Fig. 64B); rows composed of punctures larger and deeper than those on pronotum (Fig. 64A); sutural stria fine, reaching from apex to ca. apical 3/5 of the elytral length. Metathoracic wings fully developed. Mesoventrite with one distinct excavation between median carina and transverse carina (Fig. 64E); median carina of mesoventrite low (Fig. 64E); metaventrite sexually dimorphic. Legs showing distinct sexual dimorphism on protarsi, mesotarsi, metafemora, metatibiae, and metatarsi; protibiae gradually widening from base towards apex at internal margins (Figs. 65E, 65F).

Male. Middle portion of metaventrite moderate pubescent (Fig. 64F); tarsomeres 2–4 of protarsi and mesotarsi a little expanded (Fig. 65A); metafemur very slender, weakly and triangularly protuberant at about middle of posterior margins (Fig. 65C), with large dorsal projection posteroapically (Fig. 65G); metatibiae slender and feebly curved (Fig. 65C); tarsomere 1 of metatarsi very long and feebly curved (Fig. 65C); abdominal sternite 8 strongly curved (Fig. 66D); aedeagus (Figs. 66A, 66B) relatively small, its length less than 1/5 of body length; inner sac as shown in Fig. 66C.

Female. Middle portion of metaventrite sparsely pubescent mainly on basal half and almost glabrous near metacoxae (Fig. 64G); protarsi and mesotarsi slender (Fig. 65B); metafemur moderately slender (Fig. 65D), with a moderate dorsal projection posteroapically (Fig. 65H); metatibiae almost straight (Fig. 65D); tarsomere 1 of metatarsi almost straight (Fig. 65D); abdominal sternite 8 with spiculum ventrale at central point of anterior margin (Fig. 66E); coxites and stylus as shown in Fig. 66F.

Differential diagnosis. Leiodes okawai is similar to the European L. cinnamomea (Panzer, 1793) in having the male metafemora with a large dorsal projection posteroapically but can be distinguished from the latter by having the pronotum feebly sinuate at the posterior margin, the male metatibia feebly curved (Fig. 65C), and the aedeagus relatively robust (Fig. 66A). In contrast, L. cinnamomea has the pronotum straight at the posterior margin, the male metatibia strongly curved, and the aedeagus relatively slender.

Distribution. Japan: Honshu, Shikoku, and Kyushu. New to Shikoku and Kyushu.