11. Leiodes longitarsis Baranowski, 1993

Japanese name: Tsumenaga-ô-tamakinokomushi (Figs. 36–38, 113)

Leiodes longitarsis Baranowski, 1993: 43 .

Type locality. Canada, Alberta, Banff National Park.

Material examined. NORTH CHISHIMA ISLANDS: 7♂♂, 4♀♀, Ushishir, Yankicha Island, Environs of Kraternaya Bay, 20.viii.1995 (HUMS) ; 2♀♀, Simushir Island, Inland costal margin of Malaya Bay, 18.viii.1995 (HUMS) ;

1 ♂, Simushir Island, Inland costal margin of Srednaya Bay, 21.viii.1995 (HUMS). JAPAN: HOKKAIDO: 1 ♂, Rishiri Is., Mt. Rishiri, 4.–7.vii.1979, H. Henmi leg. (HUMS) ; 1 ♂, Rishiri Is., Mt. Rishiri, 8.viii.1990, K. Kishimoto leg. (FUFJ) ; 4 ♂♂, 3 ♀♀, Rishiri Is., Mt. Rishiri (alt. 1700 m), 17.–31.vii.2001, S. Hori & M. Maruyama leg. (FUFJ) .

Diagnosis. Coloration. Dorsum usually unicolor, dark reddish brown; antennae brownish; antennomeres 7, 9, 10, and basal 3/5 of antennomere 11 often darker than others; apical 2/5 of antennomere 11 often light brown.

Body 3.5–4.5 mm long, ca. 1.7× as long as wide (Fig. 36A); head densely and minutely punctate, bearing some large punctures (Fig. 36A); antennomeres 1–3 each longer than wide; antennomeres 4, 5 and 11 about as long as wide; remaining antennomeres each wider than long; antennomere 11 oval (Fig. 36C). Pronotum feebly sinuate at posterior margin, densely and minutely punctate (Fig. 36A). Elytra not transversely strigose; each elytron bearing nine rows of punctures with small number of large punctures and moderate number of fine punctures between rows (Fig. 36D); row 9 invisible in dorsal view, subhumeral row as long as ca. 1/4 or 1/3 of elytral length (Fig. 36B); elytral rows composed of larger and deeper punctures than those of pronotum (Fig. 36A); sutural stria fine, arising from apex to ca. apical half of the elytral length. Metathoracic wings fully developed. Mesoventrite without distinct excavation between median carina and transverse carina (Fig. 36E); median carina of mesoventrite low (Fig. 36E); metaventrite showing sexual dimorphism. Legs showing distinct sexual dimorphism of protarsi, mesotarsi, metafemora, metatibiae, and metatarsi; protibiae gradually and distinctly widening from base towards apex (Figs. 37E, 37F); metafemur with moderate dorsal posteroapical projection (Figs. 37G, 37H).

Male. Tarsomeres 2–4 of protarsi and mesotarsi expanded (Fig. 37A); metafemora triangularly protuberant ca. at basal 1/3 of posterior margins (Fig. 37C); metatibiae relatively slender and weakly curved inwards (Fig. 37C); metatarsi clearly longer than those of female (Fig. 37C); metaventrite with dense golden-yellow pubescence on middle portion (Fig. 36F); abdominal sternite 8 strongly curved (Fig. 38C); aedeagus as shown in Figs. 38A, B.

Female. Protarsi and mesotarsi slender (Fig. 37B); metafemora almost straight at posterior margins (Fig. 37D); metatibiae relatively robust and almost straight (Fig. 37D); metaventrite with sparse semitransparent pubescence at middle portion (Fig. 36G); abdominal sternite 8 with spiculum ventrale at central point of anterior margin (Fig. 38D); coxites and stylus as shown in Fig. 38E.

Differential diagnosis. Leiodes longitarsalis is similar to L. obesa (Schmidt, 1841) in elytral appearance, but can be distinguished from the latter by having a large body (3.5–4.5 mm), dark reddish dorsum, weakly curved male metatibiae (Fig. 37C), and the median lobe of the aedeagus with the feebly excised apex (Fig. 38A). In contrast, L. obesa has a relatively small body (3.0–4.0 mm), usually brown dorsum, relatively strongly curved male metatibiae (Figs. 83C, 83D), and median lobe with a simply rounded apex (Fig. 84A).

Distribution. U.S. A (Arizona and Colorado), Canada (Alberta, British Columbia, and Manitoba) (BARANOWSKI 1993), North Chishima Islands, Japan: Hokkaido (Rishiri Is.). New to North Chishima Islands and Japan. Leiodes longitarsis is recorded here for the first time from the Palaearctic Region.