Key to females of Chakra of India

1. Eyes small (Figures 1 (c), (16c) and 21(c), IOS at least 2× eye length (Figures 1 (b), 16(b) and 21(b)); OOL>3× MOD (Figures 1 (d), 16(d), 21(d), 23(a, i, m)) ................................... 2

- Eyes large (Figures 3 (c), 4(d), 6(c), 8(c), 10(c), 11(b), 12(c), 18(d) and 20(c)), IOS at most 1.4× eye length (Figures 3 (b), 4(b), 6(b), 8(b), 10(b), 11(b), 12(f), 18(b) and 20(b)); OOL <2.5× MOD (Figures 23 (b–h) and (23j–k)).................................................................................. 5

2. Brachypterous; T3 smooth with dense setigerous punctae; T1 unevenly foveate interspersed with short furrows posteriorly; costae on T2 closely spaced (Figure (16a)) ................................................. ................................................ Chakra parviocula sp.. nov.

- Wings fully formed (Figures 1 (e) and 21(e)); T3 either partially or entirely longitudinally striate; T1 longitudinally costate; costae on T2 spaced apart (Figures 1 (a) and 21(a)).......................................................................................................................................................... 3

3. OOL longer than POL; OOL at least 4.4× MOD (Figure 23 (a)); interantennal process triangular with a wide base (Figure 1 (b))................. Chakra agathachristieae sp. nov.

- POL and OOL subequal in length; OOL at most 3.2× MOD (Figure 23 (m)); interantennal process rectangular and elongate, gradually narrowing towards distal half (Figure 21 (b)).......................................................................................................................................................... 4

4. T3 weakly longitudinally striate (Figure 21 (a)); setose protuberances on dorsal frons, mesoscutum and mesoscutellum spaced apart (Figure 21 (a)); T1 with two short oblique carinae sublaterally on anterior margin (Figures 21 (a, d)); posterior margin of mesoscutellum weakly protruding posteriorly (Figures 21 (a, d)); mesepimeral area wider than netrion sulcus (Figure 21 (c)); posterior margin of fore wing straight (Figure 21 (e))............................................ ........................................... Chakra valluvari sp. nov.

- T3 smooth with sparse short carinae sublaterally on anterior margin; setose protuberances on dorsal frons, mesoscutum and mesoscutellum closely spaced; T1 without short oblique carinae sublaterally on anterior margin; posterior margin of mesoscutellum evenly convex; mesepimeral area and netrion sulcus subequal in width; posterior margin of fore wing angular.............................................................................................. ..................................................................... Chakra sarvatra Rajmohana and Veenakumari

5. Metascutellum rectangular with a medial notch (Figures 11 (a, d), 18(a, d), 12(a) and 14 (a, d)). ......................................................................................................................................................... 6

- Metascutellum triangular (Figures 3 (a, d), 4(a, d), 6(a, d), 8(a, d), 10(a, d) and 20(a, d)) ...................................................................................................................................................................... 8

6. Brachypterous with no marginal cilia on posterior margin of fore wing proximally (Figure 11 (e)); OOL at least 2.4× MOD (Figure 23 (g)); IOS>1.3× eye length (Figure 11 (b)); lateral pronotal area with transverse carinae posteroventrally; pronotal cervical sulcus not foveate (Figure 11 (c)); interantennal process triangular (Figure 11 (b)) ......... .................................................................................................................... Chakra kambani sp. nov.

- Wings fully formed, with marginal cilia on posterior margin of fore wing proximally (Figures 12 (h, i), 14(e) and 18(e)); OOL at most 0.7× MOD (Figures 23 (j, h)); IOS subequal to eye length (Figures 14 (b), 18(b)); lateral pronotal area posteriorly with sparse oblique carinae; pronotal cervical sulcus foveate (Figures 12 (c), 14(c) and 18(c)); interantennal process elongate and rectangular (Figures 12 (b) and 18(b)).... 7

7. Metascutellum 8× as wide as long (Figure 18 (a)); metasoma elongate,>3.5× as long as wide (Figure 18 (a)); fore wing narrow,>3.5× as long as wide (Figure 18 (e)); foveae of paracoxal sulcus round (Figure 18 (c)); metascutellum rugose (Figure 18 (a)); frons with transverse striae above central keel (Figure 18 (b))........ ....... Chakra pillaiyar sp. nov.

- Metascutellum 2× as wide as long (Figures 12 (a) and 14(a, d)); metasoma wide, at most 2× as long as wide (Figures 12 (a) and 14(a)); fore wing wide, at most 3× as long as wide (Figures 12 (h), 14(e)); foveae of paracoxal sulcus oblong (Figures 12 (c) and 14(c)); metascutellum weakly foveate (Figures 12 (a) and 14(a)); frons with transverse striae on either side of central keel (Figures 12 (f) and 14(b) .................................................. ........................................................................................................ Chakra pachmarhica (Sharma)

8. OOL at least 1.4× LOL (Figure 10 (d)); lower frons with continuous, unbroken semicircular carinae (Figure 10 (b)); horn on T1 smooth (Figure 10 (a)); four rows of robust bristles present on medial frons, continuing on to interantennal process (Figure 10 (b)); femoral depression smooth (Figure 10 (c))......................... Chakra juturna sp. nov.

- LOL either subequal or longer than OOL (Figures 23 (b–e, k)); lower frons at most with short interrupted carinae (Figures 3 (b), 4(b), 6(b), 8(b) and 20(b)); horn on T1 absent, if present longitudinally costate (Figures 3 (a), 4(a), 6(a), 8(a) and 20(a)); at most two rows of thin setae present on medial frons and interantennal process (Figures 3 (b), 4(b), 6(b), 8(b) and 20(b)); femoral depression obliquely carinate (Figures 3 (c), 4(c), 6(c), 8(c) and 20(c))............................................................................................................................... 9

9. OOL subequal to LOL (Figure 23 (e)); fore wing at least 3.5× as long as wide (Figure 8 (c)); T1 with a short horn (Figure 8 (a)); setose protuberances on head and mesonotum sparse (Figures 8 (d) and 23(e)); metascutellar spine elongate and extending over the horn on T1 (Figures 8 (a, d)) ......................................................... Chakra gotamiae sp. nov.

- OOL shorter than LOL (Figures 23 (b–d, k)); fore wing at most 3× as long as wide (Figures 3 (e), 4(e), 6(e) and 20(e)); T1 without horn (Figures 3 (a), 4(a), 6(a) and 20(a)); setose protuberances on head and mesonotum dense (Figures 3 (b), 4(b), 6(b), 20(b) and 23(b–d, k)); metascutellar spine short, not reaching T1 (Figures 3 (a), 4(a), 6(a) and 20(a))........................................................................................................................................................ 10

10. Posterior propodeal projections directed posteriorly and extending beyond anterior margin of T1 (Figures 4 (d) and 20(d)); metascutellar spine at least 1.9× as wide as long (Figures 4 (d) and 20(d)); metasoma> 2 × as long as wide (Figures 4 (a) and 20(a)); A2 and A3 subequal in length (Figures 4 (f) and 20(f)). .............................................................. 11

- Posterior propodeal projections directed laterally and not reaching anterior margin of T1 (Figures 3 (d) and 6(d)); metascutellar spine at most 1.5× as wide as long (Figures 3 (d) and 6(d)); metasoma <2× as long as wide (Figures 3 (a) and 6(a)); A3 shorter than A2 (Figures 3 (f) and 6(f)) ....................................................................................... 12

11. Femoral depression dorsally smooth and ventrally obliquely carinate (Figure 4 (c)); mesepimeral sulcus not foveate (Figure 4 (c)); central keel on frons absent (Figure 4 (b)); T4 at least 3.4× as wide as long; habitus orange-yellow (Figure 4 (a))........................ ...................................................................................................................... Chakra bournei sp. nov.

- Femoral depression entirely obliquely carinate (Figure 20 (c)); mesepimeral sulcus foveate dorsally (Figure 20 (c)); central keel on frons present (Figure 20 (b)); T4 at most 2.7× as wide as long; habitus dark brown (Figure 20 (a))............................................... ......................................................................................................... Chakra sanghamittae sp. nov.

12. Central keel absent (Figure 6 (b)); lower frons with short transverse and oblique carinae on either side of imaginary central keel (Figure 6 (b)); oblique carinae on ventral femoral depression sparse (Figure 6 (c)); basal foveae of T2 3× as long as wide (Figure 6 (a)) .............................................................................. Chakra galathea sp. nov.

- Central keel present (Figure 3 (b)); lower frons with weak transverse and oblique reticulations on either side of central keel (Figure 3 (b)); oblique carinae on ventral femoral depression dense (Figure 3 (c)); basal foveae of T2 at most 2× as long as wide (Figure 3 (a)) ........................................... ........................................... Chakra alexandra sp. nov.