Leucophenga quadripunctata (de Meijere, 1908)

(Figs 2 H, 6I, 6J, 10I, 10J, 14C, 14D, 24)

Drosophila quadripunctata de Meijere, 1908: 154; Okada, 1990: 683.

Diagnosis. Wing R2+3 vein with a short branch subapically (Fig. 2 H); paramere deeply concave mediodorsally (Fig. 24 C). This species is similar to L. cornuta sp. nov. in the wing R2+3 vein with a short branch subapically (see lower that species).

Description. Ocellar triangle brown, with 3–7 setae above ocellar setae. Frons yellowish brown to brown. Pedicel yellowish brown; first flagellomere yellow. Clypeus brown. Palpus brownish yellow. Mesonotum nearly yellowish brown, slightly brown before scutellum (Fig. 6 I). Postpronotal lobe yellow (Fig. 6 J). Acrostichal setulae in 13–15 irregular rows (Fig. 6 I). Scutellum yellowish brown, dark brown at tip and pale around apical scutellar setae (Fig. 6 I). Pleura with an unapparent, yellowish brown, longitudinal stripe above (Fig. 6 J). Katepisternum and mesopleuron mostly brownish (Fig. 6 J). Wing (Fig. 2 H): Costal vein between R2+3 and R4+5 distally with 5–8 peglike spinules on ventral surface. Halter slightly brownish apically (Fig. 6 J). Abdominal tergites brown to black, with yellow patches (Fig. 10 I, J). Male terminalia: Epandrium with pubescence near posterior and ventral margins and 6 setae near posterior margin per side (Fig. 24 A). Paramere with pubescence and 4 sensilla subapically (Fig. 24 C). Aedeagus pubescent apically (Fig. 24 D).

Measurements. BL = 3.90–4.67 mm in ♂, 3.43–4.13 mm in ♀ (range in 5♂ and 4♀), THL = 1.90–2.07 mm in ♂, 1.53–1.97 mm in ♀, WL = 3.00– 3.60 mm in ♂, 2.93–3.57 mm in ♀, WW = 1.42–1.67 mm in ♂, 1.37–1.60 mm in ♀, arb = 6–8/3–4, avd = 0.73–0.96, adf = 1.33–1.94, flw = 1.57–2.00, FW/HW = 0.36–0.40, ch/o = 0.03–0.04, prorb = 0.51–0.75, rcorb = 0.51–0.82, vb = 0.30–0.44, dcl = 0.44–0.60, presctl = 0.49–0.67, sctl = 1.12–1.63, sterno = 0.74–1.00, orbito = 1.64–2.56, dcp = 0.21–0.34, sctlp = 0.97–1.27, C = 1.63–2.06, 4c = 1.41–1.94, 4v = 1.64–2.36, 5x = 0.95–1.37, ac = 2.70–3.44, M = 0.51–0.74, C3F = 0.86–0.92.

Specimens examined. CHINA: 1♀ (SCAU, No. 123112), Dabang, Jiayi, Taiwan, 800m, 15.x.2012, ex tree trunk, HW Chen; 4♂, 2♀ (SCAU, Nos 123113 –18), Maoershan, Guilin, Guangxi, 700m, 23.ix.2006, ex tree trunks, JJ Jiang; 1♂, 1♀ (SCAU, Nos 123119, 20), Jiaoye Park, Kunming, Yunnan, 24°32'N, 101°01'E, 2100m, 6.iii.2001, ex tree trunks, JJ Gao; 1♀ (SCAU, No. 123121), Weixi, Yunnan, 1900m, 29.vii.2004, ex tree trunk, HW Chen.

JAPAN: 4♂ (SCAU, Nos 122946, 123122, 23), Minamiohsawa, Hachioji, Tokyo, 35˚37'N, 139˚23'E, 130m, 21.vi.2002, ex tree trunks, HW Chen.

Distribution. China (Taiwan, Guangxi, Yunnan), Russia (Far East), Japan (Honshu), Korea, Thailand, Sri Lanka, Malaysia (Sarawak, Kuala Lumpur), Indonesia (Java, Sumartra), Papua New Guinea (Wau), Australia (Queensland).