Parastenolechia Kanazawa

Parastenolechia Kanazawa, 1985: 6 .

Type species: Parastenolechia asymmetrica Kanazawa, 1985 . Origo Omelko, 1988: 156 .

Type species: Telphusa argobathra Meyrick, 1935, by original designation. Laris Omelko, 1988: 152 . New Synonymy

Type species: Laris collucata Omelko, 1988 .

Description. Imago. Labial palpus with third and second segments subequal in length. Clypeus with ventral margin sinuate mesially. Antenna almost filiform but slightly serrate distally, about 3/5 forewing length, shorter and thicker in male than female. Ocellus absent. Posterior area of sitophore with four campaniform sensilla in line except one offset on right side; anterior area with four campaniform sensilla. Forewing (length/ width ratio 4.0) with raised scale tufts, R5 and M1 separate, M2 and M3 connate, CuA1 absent, CuA2 almost vestigial (Fig. 12); median fascia directed from apex of costa toward posterior margin. Hindwing (length/width ratio 5.0) with M1 absent, M2, M3, and CuA1 separate. Male abdomen with tergum VIII considerably reduced; sternum VIII well developed, without posterior or anterolateral hair pencils.

Male genitalia: uncus flattened, slightly concave at middle of apex; gnathos almost as long as uncus, with small, cushionlike, mesially projected apex; tegumen narrower mesially than basally, strongly widened anteriorly; valva asymmetrical, divided into flagelliform costa, longer than length of tegumen, and broad sacculus; tegumen basal width/length ratio 0.9; vinculum absent; saccus fused with ventrobasal part of slender phallus; phallus slightly shorter than tegumen, fulcrum well developed; no cornuti.

Female genitalia: apophyses anteriores thick and slightly curved dorsally; apophyses posteriores elongate, 2.0–2.6 x as long as apophyses anteriores; ostium bursae on intersegmental membrane posterior to sternum VII; antrum membranous; signum a semiovate plate with pair of anterolateral lobes.

Larva. 5.5–6.5 x 1.2–1.5 mm, head width 0.67 mm; body compact, reddish brown, with posterior areas of abdominal segments bluish gray, giving a transversely banded appearance; pinnacula inconspicuous; head brownish black, with broadly incised labrum, antenna longer than in Stenolechia gemmella; prothoracic shield brownish black, pale on anterior and ventral margins, narrowly divided into two plates anteriorly, widely divided posteriorly; anal shields and plates of anal prolegs brownish black; abdominal prolegs short, crochets uniordinal; anal comb reduced to small spines; prothorax with only L1 and L2 present; all setae on meso- and metathorax on isolated pinacula; Abdomen with microscopic, conical granulations dorsally; L1 and L2 horizontal on A1–8, transverse on A9; A9 and A10 with enlarged pinacula, all setae isolated except L group (Patoèka 1977).

Pupa. Maxillary palpi separated from genae; contiguous boundary between maxillary palpi and mesothoracic legs short; frons lacking a tubercle; frontoclypeal suture almost straight; prothoracic legs adjacent to oculi; abdominal segment VII not edged caudally by setae (Patoèka and Turcáni 2005).

Diagnosis. Parastenolechia species are similar to those of Stenolechiodes in the wing pattern and male genitalia, but differ by having M1 vein absent in hindwing and a signum present in female genitalia.

Hosts. Fagaceae: Quercus pubescens Willd. ( P. nigrinotella). (Patoèka 1977).

Diversity and distribution. The twelve species of Parastenolechia occur in Europe and eastern Asia, with highest diversity in the latter region (Elsner 1995 [1996]; Huemer and Karsholt 1999; Kanazawa 1991; Okada 1961; Omelko 1988; Park 1993, 2000b; Park and Ponomarenko 2006).

Notes. Laris was described to include two species in two subgenera and was differentiated from other genera by a strongly differentiated sacculus (Omelko 1988). Park (1993) synonymized subgenus Origo with Parastenolechia and differentiated the nominal subgenus, including L. collucata, from Parastenolechia based on the venation of forewing with CuA2 present (absent in Parastenolechia), R4 and R5 with a long stalk, and male genitalia with lobate distal portion of the valval sacculus. However, the forewing has CuA2 present in both genera, although it is almost vestigial in Parastenolechia (Kanazawa 1985), and the length of R4+5 varies among species in other genera of Teleiodini . The valval sacculus is distally lobate in both L. collucata and species of Parastenolechia, as illustrated by Kanazawa (1985). Species of Parastenolechia and L. collucata are the only taxa of Teleiodini with a forewing lacking CuA1. In addition they have an uncus with a truncate apex and a gnathos with the ventral part being cushionlike with a median projection, which are synapomorphic for the polytomy of Parastenolechia, Stenolechia, and Chorivalva (Lee 2007) . Therefore, Laris is regared as a junior synonym of Parastenolechia .