Genus Anchonidium Bedel, 1884

Rostrum with long and straight scrobes, these flattened and conjoint at underside towards base (Figs 1 A-B). 2/3 of underside of rostrum from tip on bearded (Figs 1B, D, G), rather densely covered by longer yellowish bowed bristles, dorsum of rostrum flattened, tricarinate (Fig. 1A), angulate towards apex and base (Fig. 1D). First club segment several times longer than following ones, cup-shaped; antennal funiculus with 7 antennomeres, longest first segment, second one half of the length and following ones transverse, steady broadened towards club (Fig. 1I). Pronotum roundish, characteristically sculptured with a V-shaped impression on upper side, and a longitudinal carina at disc, strongly constricted before fore margin, strongly punctuate (Fig. 1F); Procoxae almost touching each other, separated by thin interspace (Fig. 1C); Mesocoxae separated by half their diameter; hind margin of metasternum with sharp pointed edges sidewards (Fig. 1E). Metacoxae separated by 1.5 their diameter; in middle of metacoxae on first ventrite coarsely punctuate with especially large, roundish-oval punctures or confluent huge punctures in the shape of an oval groove (Fig. 1J); elytra with striae coarsely punctuate, intervals reduced to narrow elevated ridges between stria, uneven intervals higher elevated (Fig. 2 D-F); vestiture of body and legs dominated by yellowish bowed bristles, these denser standing along sides and underside of rostrum, on pronotum, on femora and tibiae, and along uneven ridges on elytra; Femora unarmed; Tibiae all uncinate at outer angle (Fig. 1H), third tarsal segment bilobed, claws simple, free; genital organs: penis tube shaped in cross-section and bowed in lateral view (Figs 3 I-L), bursal atrium strongly sclerotized (Fig. 4J, K).

Remark: One of the most apparent differences between the three species of Anchonidium in the narrow sense and the others, is the very deviating shape of the penis of A. caucasicum, which is flat in cross-section (and not tube shaped as in Anchonidium sensu stricto) with its tip cut in middle (Fig. 3 O-P).

Diagnostic key to species of the genus Anchonidium

1A Body without yellowish bowed bristles only sparse setae on elytra; elytra oval, laterally evenly rounded; pronotum without sculpturation, evenly rounded at disc; penis flat in cross section, characteristically cut in middle at apex (Fig. 3 O-P), species from Caucasian region...................................................................... Anchonidium sensu lato

1B Body with yellowish bowed bristles, dense on rostrum, pronotum, along ridges on elytra, and on legs; elytra laterally more parallel; pronotum strongly sculptured (Fig. 1F); penis tube-shaped in cross section, apex rounded, blunt or sinuate (Figs 3 A-L), species from western Europe and northern Africa ..... Anchonidium sensu stricto, 2

2A Ventral side of pronotum at fore margin with a faint and shallow rostral notch or channel (Fig. 1C). First ventrite with oval groove between metacoxae (Fig. 1J); elytra laterally rounded, shorter, 1.4-1.5 times as long as wide; 10 to 12 punctures on elytral disc from base to declivity; widest in first third (Fig. 2 D-F); penis thin (lateral view), slender with almost rectangular apex and rounded sides (dorsal/ventral view; Fig. 3 I-L). Great Britain (Cornwall), W- and NW-France, N-, NW- and S-Spain, N-Portugal, N-Morocco ............................................... A. unguiculare

2B Ventral side of pronotum at fore margin only sinuate, no rostral notch or channel detectable (Fig. 2B). First ventrite without groove (Fig. 2B), but very coarse and/or oval punctures between metacoxae; elytra parallel sided, longer, 1.5-1.8 times as long as wide; 14 to 16 punctures on elytral disc from base to declivity, widest in or behind middle (Fig. 2A, C); penis short, blunt and thick in lateral view (Figs 3 A-H)..................................................................... 3

3A Elytra widest along middle (Fig. 2A), 1.6-1.8 times as long as wide; tip of penis broad, margin rounded (Fig. 3 E- H). Portugal (Serra de Monchique) .......................................................................................... A. braunerti sp. nov.

3B Elytra widest behind middle (Fig. 2C), 1.5-1.6 times as long as wide; tip of penis very broad, spatula-shaped with sinuate fore margin (Fig. 3 A-D). Portugal (Serra da Estrela, Serra do Marão)................... A. spathiferum sp. nov.

Remarks: As both new species of the genus Anchonidium are – based on their external morphology – very similar to each other and share in general most characters with A. unguiculare, except for those ones mentioned in the key below, the descriptions of the new species are provided in a condensed form.