Cicindela (s. str.) coerulea shantungensis Mandl, 1970

Figs 55, 56, 134, 136, 150–154, 171–195

Cicindela coerulea shantungensis Mandl, 1970: 72, Abb. 3 figs 3–5 (Type locality— China, Tsingtau). Cicindela tricolor M.F. Adams, 1817: Grum-Grzhymailo 1899: 240. Cicindela hybrida nitida Lichtenstein, 1796: Wu 1937: 15; Institute of Zoology Academia Sinica 1986: 417; Hu 1992: 374; Zhu et al. 1999: 26. Cicindela coerulea Pallas, 1773: An 2011: 276, fig. 7, 277. Cicindela coerulea nitida Lichtenstein, 1796: Kim 1998: 354; Werner 1991: 18, T. 14 fig. 103; Kim et al. 2005: 6, fig. 6. Cicindela coerulea shantungensis Mandl, 1970: Mandl 1981: 28; Cassola & van Nidek 1984: 10; Wiesner 1992: 118; 2020: 184; Lorenz 1998: 43; 2005: 131; Puchkov & Matalin 2003: 105; Shook & Wiesner 2006: 10; Wu & Shook 2010: 78; Wu 2011: 24; Putchkov & Matalin 2017: 227.

Type material. Paratypes of C. coerulea shantungensis Mandl, 1972: 1 ♀ (Figs 171, 172)— China, Kiautschou [handwritten white label], Paratypus [typed red label], Cicindela coerulea ( nitida) shantungensis m., det. K. Mandl 1970 [handwritten and typed white label], coll. K. Mandl [typed light-blue label]; NHMW-ZOO-COLL, 0002628 [printed white label]; 1♀ (Figs 152, 173, 174, 178)— China, Kiautschou [handwritten white label], Paratypus [typed red label], Cicindela coerulea ( nitida) shantungensis m., det. K. Mandl 1970 [handwritten and typed white label], coll. K. Mandl [typed light-blue label], 222, Cicindela coerulea ( nitida) shantungensis Mandl [handwritten white label]; NHMW-ZOO-COLL, 0002627 [printed white label] (both NHMW).

Additional material. CHINA: Liaoning Province: 4♂♂ 3♀♀ — Huludao City, Jianchang County, Yangmadianzi Township, 40.53912°N 119.78307°E, 15.VII.2022, leg. Xuekui Wang (cXX) ; Hebei Province: 1♂ — China, Can zhou (= Cangzhou), 1500 m, VI.1991, leg. J.M. Bousquet (cJW); 3♂♂ 2♀♀ — Qinhuangdao City, Haigang District, Duzhuang Town, Xiaojiagang Village, 40.019537°N 119.558324°E, 19. V.2023, leg. Zhiyong Bai (cXX) ; Beijing Municipality: 3♂♂ 1♀ — Nord Pekin, Jehol, A. David, 1865 (MNHN) ; 2♂♂ — Peking (ZIN); Shanxi Province: 1♀ — Shan-si, 18. VI.1884, G. Potanin (ZIN); Shandong Province: 1♂ —1898, China, Kiautschou (NHMW); 1♀ — Tsingtau (HNHM) ; 2 ex. — ibid, S. Glaue; 2 ex. — ibid, 31.VIII.1919, E.S. Märker (both MNHUB); Gansu Province: 1♂ —sanads Pin-Faun-Cha, 8.IV.[18]90, Grum [- Grzhymailo] (ZIN); 1♂ — Kansou or., K’ing-iang-fou (ZMUM) ; Qinghai Province: 1♂ 3♀♀ —canyon of Moudjik (= Si-He) River, 3–7.VII.[18]90, Grum [- Grzhymailo] (ZIN); WITHOUT EXACT LOCALTY : 1♀ — Collect. Konschegg (ZMUM) .

References. CHINA, Beijing Municipality: Peking (Mandl 1951); Peking, 39 o 55’N 116 o 25’E (Mandl 1981); Beijing, Yanqing District, 40.38684 N 115.86325 E, 09.X.2011, Terry Townshend (https://www.inaturalist. org/observations/50020234); Beijing, 40.64235 N 117.46375 E, 22.V.2019, Shan Gui, (https://www.inaturalist. org/observations/27913762); ibid, 40.02167 N 116.14653 E, 40.06845 N 116.1495 E, 40.14761 N 116.0647 E, 40.14709 N 116.08837 E, 40.10333 N 116.0267 E, 40.12321 N 116.13432 E, VI.2023, Zhongqi Wang (https:// www.inaturalist.org/observations/188438646); Beijing, Mentougou District, 40.06896 N 116.02095 E, 02.VI.2023, Yihang Li (https://www.inaturalist.org/observations/167686381); Shanxi Province: Mien Shan, 3700, 37 oN 112 o 30’E (Mandl 1981); Shandong Province: Tsingtau, Kiautschou (Mandl 1970); Qinghai Province: Hainan Tibetan Autonomous Prefecture, Guinan County, 35.68101 N 101.060027 E, 08.07.2016 (https://www.inaturalist. org/observations/69748970); Hainan Tibetan Autonomous Prefecture, Xinghai County, 36.04694 N 100.11991 E, 13.IX.2020 (https://www.inaturalist.org/observations/62014051); ibid, 36.223347 N 100.11401 E, 22.08.2021 (https://www.inaturalist.org/observations/92649596); Gansu Province: Jingyuan County, Baiyin City, 36.94422 N 105.15422 E, 05.VII.2024 (https://www.gbif.org/occurrence/4908088997); SOUTH KOREA: Seoraksan Mountain Natural Reserve; Jeju-do, Hallasan Mountain Natural Reserve (both An 2011); Gyehwado, Gimje-gun (Kim et al. 2005); 36.7 N 128.1 E, 16.07.2013 (https://www.gbif.org/occurrence/3444542042); 36.7 N 126.6 E, 16.05.2019 (https://www.gbif.org/occurrence/2825849468); Henan Province: Jiyuan City, Sanmenxia City (Lingbao), Luoyang City (Yanshi District and Luoning County), Zhengzhou City, Kaifeng City, Zhumadian City (Xiping County), Nanyang City, Xinyang City (Gushi County) (all Zhu et al. 1999).

Diagnosis. Cicindela coerulea shantungensis differs from other subspecies by a reduced and relatively narrow white elytral pattern (HI = 5.0–9.0, mean = 7.3) with humeral lunula divided on two isolated spots and oblique branch of middle band without rounded apical extension (Figs 171, 173, 175–189), by relatively long and thin basal portion of aedeagus (Figs 55, 56), and by proportion of pronotum (Figs 150–154, 244).

Redescription. TL = 13.5–15.2 mm (mean = 14.69 mm, n = 7) in males, 15.2–16.1 mm (mean = 15.71 mm, n = 7) in females (Figs 171, 173, 175–177).

Head blue or bluish-green, genae bright blue or green, clypeus, supraorbital plates, central portion of frons, vertex and occiput cupreous with golden tinge. Labrum with 6–13 (mean = 8, n = 15) submarginal setae, LW/LL = 2.0–2.3 (mean = 2.13, n = 7) in males, 1.77–2.23 (mean = 1.97. n = 7) in females (Figs 134, 136). Mandibles relatively thin, ML/MW = 6.0–6.33 (mean = 6.17, n = 3) in males, 6.0–8.0 (mean = 7.07, n = 3) in females. Scape besides 3–4 apical setae with a single, rarely with 2–4 additional setae, in some specimens glabrous.

Pronotum green or bluish-green with golden-cupreous reflection especially in central portion of anterior and posterior lobes, deep anterior and posterior grooves as well as thin midline bright blue; PW/PL = 1.21–1.37 (mean = 1.32, n = 15) in males, 1.32–1.45 (mean = 1.38, n = 12) in females (Figs 150–154). Thoracic segments blue or bluish-green, in some specimens all epimerae and episternae green or blue-green with light golden lustre.

Elytra cupreous often with golden lustre or purple-cupreous, with distinct granulation most developed on humeri especially in females; EL/EW = 1.4–1.7 (mean = 1.57, n = 27); white elytral pattern as a rule relatively narrow, humeral lunula divided on two isolated spots, rarely with virtually invisible medial portion, middle band in most specimens with short or long thin oblique branch without or rarely with slightly rounded apical extension; marginal band undeveloped, rarely only marked; apical lunule complete with thin medial portion (Figs 171, 173, 175–189).

Abdominal sternites blue or greenish-blue with light golden lustre.

Aedeagus with relatively long and thin basal portion and small stout apical hook (Figs 55, 56), EL/AL = 1.65–2.0 (mean = 1.8, n = 10). Internal sac relatively short but rounded towards to ventral side so В contact with the aedeagus tube (Figs 190–195).

Distribution. CHINA: Beijing Municipality, Liaoning, Hebei, Shandong, Shanxi, Henan, Shaanxi, Gansu, Qinghai Provinces; SOUTH KOREA: Gangwon and Jeonbuk States, South Chungcheong, North Gyeongsang and Jeju Provinces (Fig. 246).