Cicindela (s. str.) coerulea grumi Tschitschérine, 1903

Figs. 63, 64, 138, 140–144, 198–202, 207–214, 221–232, 234–236, 238–240

Cicindela przewalskii var. grumi Tschitschérine, 1903: 98 (Type locality—Mons Nan-Shan: region do la riv. Babo-che (= EdzinGol), 11 000’–12 700’ d’altitude). = Cicindela przewalskii var. kozlovi Tschitschérine, 1903: 99(Type locality—Thibet or.: region Dabasun-gobi (au SE du lac Dalaï-dabasu et au N de la Tamyrtyn-ula), synonymized by Schilder (1953: 564). = przevalskii var. grumi [auct.] = przevalskii var. kozlovi [auct.] = kozlowi [auct.]

Cicindela hybrida grumi Tschitschérine, 1903: Jakobson 1905: 190; Horn 1905a: 157, 158; 1905b: 27; 1915: 336; 1926: 221; 1930: 402; Winkler 1924: 4; Wu 1937: 14.

Cicindela grumi grumi Tschitschérine, 1903: Schilder 1953: 564.

Cicindela coerulea grumi Tschitschérine, 1903: Mandl, 1935: 294 fig. 11, 296 fig. 13; 1936: 13; Horn 1938: 46, Taf. 63 figs 27, 28, Taf. 64 fig. 1; Cassola & van Nidek 1984: 10; Werner, 1991: 18, T. 15 fig. 110; Wiesner 1992: 118; 2020: 184; Lorenz 1998: 42; 2005: 131; Puchkov & Matalin 2003: 104; Shook & Wiesner 2006: 10; Wu & Shook 2010: 78; Wu 2011: 24; Putchkov & Matalin 2017: 227.

Cicindela hybrida kozlovi Tschitschérine, 1903: Jakobson 1905: 190; Horn 1905a: 158; 1905b: 26; 1915: 336; 1926: 221; 1930: 401; Winkler 1924: 3; Wu 1937: 14.

Cicindela coerulea kozlovi Tschitschérine, 1903: Mandl, 1935: 294 figs 12, 13, 296 fig. 14, 300 fig. 18; 1936: 14; Horn 1938: 46, Taf. 64 fig. 2; Cassola & van Nidek 1984: 10; Werner, 1991: 18, T. 15 fig. 111; Wiesner 1992: 118; 2020: 184; Lorenz 1998: 42; 2005: 131; Puchkov & Matalin 2003: 104; Shook & Wiesner 2006: 10; Wu & Shook 2010: 78; Wu 2011: 24; Putchkov & Matalin 2017: 227.

Type material. Lectotype of Cicindela przewalskii var. grumi Tschitschérine, 1903 (designated here), ♂ (Figs 138, 198, 199, 234, 238)—р. БадО-хО и ХЫ-хО-дО, 13.VIII.90, Грум. [handwritten white label], silver circle, Type, v. grumi m., Tschitschérine det. [handwritten and typed white label], LECTOTYPUS, Cicindela przewalskii var. grumi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label] (ZIN). Paralectotypes of Cicindela przewalskii var. grumi Tschitschérine, 1903 (designated here): 2♂♂ 4♀♀ (Figs 63, 140, 207–210, 221–226)—same labelled as Lectotype, but PARALECTOTYPUS, Cicindela przewalskii var. grumi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label]; 3♂♂ —р. БадО-хО и ХЫ-хО-дО, 13.VIII.90, Гр. ГржимайлО” [handwritten white label], PARALECTOTYPUS, Cicindela przewalskii var. grumi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label]; 1♀ —р. БадО-хО и ХЫ-хО-дО, 13.VIII.90, Гр. ГржимайлО [handwritten white label], Cic. gemmata potanini Dokht., A. Semenov-Tian-Shansky det., VI.30 [handwritten and typed label], PARALECTOTYPUS, Cicindela przewalskii var. grumi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label] (all ZIN); 3 ex. —р. БадО-хО и ХЫ-хО-дО, 13.VIII.90, Гр. ГржимайлО [handwritten white label], PARALECTOTYPUS, Cicindela przewalskii var. grumi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label] (SDEI).

Lectotype of Cicindela przewalskii var. kozlovi Tschitschérine, 1903 (designated here), ♂ (Figs 141, 200, 201, 211, 227, 236, 240)—ДабасуН ГОби, ЭКсп. КОЗлОва, 1/ 2.VIII.1901 [handwritten white label], golden circle, Type, Przewalskii v. kozlovi m., Tschitschérin det. [handwritten and typed white label], LECTOTIPUS, Cicindela przewalskii var. kozlovi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label] (ZIN). Paralectotypes of Cicindela przewalskii var. kozlovi Tschitschérine, 1903 (designated here): 2♂♂ 1♀ (Figs 64, 143, 213, 228– 230)—ДабасуН ГОби, ЭКсп. КОЗлОва, 1/ 2.VIII.1901 [handwritten white label], golden circle, Type, v. Kozlovi m., Tschitschérine det. [handwritten and typed white label], PARALECTOTIPUS, Cicindela przewalskii var. kozlovi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label] (ZIN); 3 ex. — ДабасуН ГОби, ЭКсп. КОЗлОва, 2.VIII.1901 [handwritten white label], PARALECTOTIPUS, Cicindela przewalskii var. kozlovi Tschitschérine, 1903, des. A. V. Matalin, 2024 [printed red label] (SDEI).

Additional material. CHINA, Qinghai Province: 1♀ — Quinghai-Lake, eastern edge, 07.07.2011, leg. M. Schülke (cDW) ; 4♂♂ 4♀♀ — Qinghai Prov., Qinghai Lake nature reserves, east of lake, 36.83 oN 100.75 oE, 3202 m, 18.IV.2015, leg. Zhenhua Liu, Menjie Jin (SYSU) ; 3♂♂ 1♀ — China NW, Qinghai, Wulan (= Ulan), h ~ 3700–4000 m, 16– 21.07.1997, leg. A. Wrzecionko (SIZ) ; 1♂ — ibid, 1–6.VII.1995; 2♀♀ — ibid, 3500 m, 24– 25.7. 1992, leg. S. Kocman (both cJW); 2 ex. — China, Qinghai-Haixi, Wulan (= Ulan), 40 km N Chakayan Nuur ( Lake), 25.07.1992, leg. S. Kocman ; 2 ex. — ibid, 01.07.1995, leg. A. Wrzecionko (both cJG); 2 ex. — ibid, 09.07.2001 (cFB).

References. CHINA, Qinghai Province: Haibei Tibetan Autonomous Prefecture, Haiyan County, Ganzihe Township, 37.150331 N 100.540469 E, 15.07.2021, 23.07.2022, Zhang Qianyi (https://www.inaturalist.org/ observations/193423392#data_quality_assessment)

Diagnosis. This subspecies is easily distinguished from other subspecies by transverse or virtually transverse, slightly sinuate middle band often with rounded apical portion (Figs 198, 200, 202, 221–232), as well as by the proportion of labrum and pronotum (Figs 244, 245). From nominotypical subspecies and C. coerulea nitida C. coerulea grumi is clearly distinguished by short slightly rounded towards to ventral side internal sack so В uncontact with the aedeagus tube (Figs 234–236, 238–240 vs. Figs 65–70 and Figs 117–132). Moreover C. coerulea grumi significantly differs from C. coerulea przewalskii by the proportions of pronotum and labrum (Figs 207–214 vs. Figs 203–206; Figs 138, 140–144 vs. Figs 137, 139; Figs 244, 245).

Redescription. TL = 12.1–14.0 mm (mean = 13.14 mm, n = 12) in males, 12.8–15.0 mm (mean = 14.11, n = 8) in females. Most specimens unicolor brown or cupreous-brown, sometimes black, blue or green, rarely purple-cupreous (Figs 198, 200, 202).

In brown specimens genae, clypeus, antennal plates, lateral portions of frons and shallow intraocular cavities bright green with cupreous or golden-cupreous reflection, base of genae bright blue; central portion of frons and vertex as well as supraorbital plates bright cupreous, sometimes with golden-purple lustre; occiput cupreous-bronze; shallow interocular cavities bright blue or greenish-blue; in green specimens genae with light golden-cupreous tinge while clypeus and intraocular cavities with bright blue lustre; in some specimens head completely blue or black. Labrum with 6–13 (mean = 8, n = 12) submarginal setae, LW/LL = 2.09–2.33 (mean = 2.18, n = 12) in males, 2.0– 2.27 (mean = 2.16, n = 86) in females (Figs 138, 140–144). Mandibles relatively stout, ML/MW = 6.0–8.0 (mean = 7.08, n = 9) in males and 7.2–8.25 (mean = 7.63, n = 6) in female. Scape besides 3–4 apical setae glabrous, rarely with no more than two additional setae.

Pronotum purple-cupreous or cupreous-green with golden reflection, anterior and posterior grooves as well as midline bright blue, bluish-green or golden-green; in some specimens completely green, blue or black (Figs 207–214); PW/PL = 1.41–1.52 (mean 1.45, n = 13) in males and 1.43–1.57 (mean = 1.49, n = 10) in females. Proand mesothorax bluish-green with light golden tinge; metathorax blue-violet, cupreous-green laterally with distinct golden reflection; pro- and mesepimeron as well as metepisternum bright cupreous with golden-green reflection; mesepisternum green with golden-cupreous or cupreous-bronze lustre; in some specimens thoracic segments entirely black or black-brown.

Elytra brown, cupreous-bronze or purple-cupreous with dense golden, blue or greenish-blue punctures sometimes black, blue or green; EL/EW = 1.49–1.63 (mean = 1.56, n = 13) in males and 1.49–1.60 (mean 1.54, n = 10) in females; white elytral pattern complete, relatively wide; humeral and apical lunulae always isolated from slightly curved middle band with short marginal portion, in some specimens middle band lacking rounded apical extension (Figs 198, 200, 202, 221–232).

Abdominal sternites black, blue or bluish-green with light golden or golden-green tinge.

Aedeagus relatively short with relatively short and wide basal portion (Figs 63, 34), EL/AL= 1.74–1.96 (mean 1.83, n = 11). Internal sac with short slightly rounded towards to ventral side internal sack so В uncontact with the aedeagus tube (Figs 234–236, 238–240).

Distribution. CHINA: Qinghai Province (Fig. 246).

Notes. Cicindela przewalskii var. grumi and Cicindela przewalskii var. kozlovi were described from northern and southern environs of Kuko Nor (= Qinghai) Lake, respectively (Tschitschérine 1903). Re-examination of the types of both these taxa leads me to consider them conspecific. The similarity of the structure of the labrum (compare Fig. 138 with Figs 141, 142 and Fig. 140 with Figs 143, 144), pronotum (compare 207–210 with Figs 211–214) and internal sac of the aedeagus (compare Fig. 234 with Figs 235, 236 and Fig. 238 with Figs 239, 240), as well as the complete coincidence of the main proportions of the body (Figs 243–245) help justify this opinion. At the same time, both taxa clearly differ from the most related to them, C. coerulea przewalskii, by the form of pronotum (Figs 207–214 vs. Figs 203–206), white elytral pattern (Figs 221–232 vs. Figs 215–220) and the main proportions of the body (Figs 243–245). Thus, I accept the opinion of Schilder (1953: 564), who considered Cicindela kozlovi Tschitschérine, 1903 as a synonym of Cicindela grumi Tschitschérine, 1903 .