Myrmetes paykulli Kanaar, 1979

(Figs. 11, 61, 95, 126, 160–161, 491–507)

Hister piceus: PAYKULL (1809): 231, nec MARSHAM (1802): 97 (misidentification).

Saprinus piceus: ERICHSON (1839): 676; MARSEUL (1855): 505, t. XIX, Fig. 122.

Gnathoncus piceus: JACQUELIN- DUVAL (1858): 112.

Myrmetes piceus: MARSEUL (1862):515, t. XIII, Figs.1, 1a; THOMSON (1862): 243; SCHMIDT (1885a): 318; GANGLBAUER (1899): 378; REITTER (1909): 290, t. 67, Fig. 4; BICKHARDT (1916 –1917): t. IV, Fig. 35; REICHARDT (1941): 172, Fig. 82; HORION (1949): 346; MAZUR (1973): 41, Fig. 88; KRYZHANOVSKIJ & REICHARDT (1976): 122, Fig. 164.

Myrmetes paykulli Kanaar, 1979: 24 . New substitute name for Hister piceus Paykull, 1809, nec Marsham, 1802).

Myrmetes paykulli: VIENNA (1980): 113, Fig. 42; MAZUR & KASZAB (1980): 63, Fig. 32; MAZUR (1981a): 94, Fig. 128; MAZUR (1984): 107; MAZUR (1997): 217; YÉLAMOS (2002): 254, Figs. 122D, 128; MAZUR (2004): 95.

Note. Sensory structures of the antennal club were studied by DE MARZO & VIENNA (1982a).

Type locality. Sweden, near Uppsala.

Material examined. CZECH REPUBLIC: BOHEMIA: Liberec, Ruprechtice, xii.1997, nest of Formica rufa, 1 ♁, M. Švarc lgt .; Horní Jelení, 19.x.2001, 1 ♁, V. Zieris lgt. MORAVIA: Hnanice env., Na Fládnici, 30.iv.2003, 1 spec., Jiří Hájek lgt .; Srbce env., 28.xi.2001, 1 spec., V. Zieris lgt. (TLAN) .

Redescription. Body length: PEL: 1.70–1.775 mm; APW: 0.525 –0.625 mm; PPW: 1.375 – 1.425 mm; EL: 1.175–1.25 mm; EW: 1.575 –1.675 mm.

Body (Figs. 491–492) shortly oval, rounded, dorsally moderately convex, flattened ventrally, cuticle reddish-brown to dark brown with silk metallic luster; legs, mouthparts and antennae reddish.

Antennal scape (Fig. 495) slender, surface imbricate, with few short setae, club (Fig. 493) elongated, flattened, without visible articulation, entire surface with dense short sensilla intermingled with sparse somewhat longer erect sensilla; sensory structures of antennal club (Fig. 11) in form of four rather small round circular sensory areas on ventral side and one large round vesicle on internal distal margin of ventral side.

Mouthparts. Mandibles (Fig. 95) with curved outer margin, acutely pointed; sub-apical tooth on inner margin of left mandible minute; disc of labrum (Fig. 61) imbricate, flattened; labral pits tiny, single weakly sclerotized seta present in each pit; labral fold significantly less developed; terminal labial palpomere thickened, its width about half its length; mentum sub-trapezoid, anterior angles (Fig. 126) somewhat produced; anterior margin shallowly emarginate, with sparse, moderately long setae; lateral margins with one row of short ramose setae; disc of mentum glabrous; cardo of maxilla on lateral margin with few short setae; stipes triangular, with three short weakly sclerotized setae; terminal maxillary palpomere elongated, its width about one third its length, about three times as long as penultimate.

Clypeus (Fig. 494) rectangularly rounded laterally, imbricate-punctate; frontal and supraorbital striae absent; frontal disc (Fig. 495) microscopically imbricate-punctate; eyes flattened, visible from above.

Pronotum (Fig. 491) transverse, about twice as broad as long; pronotal sides strongly narrowed anteriorly; apical angles conspicuous; marginal pronotal stria weakly impressed but complete; pronotal disc smooth; pronotal hypomeron glabrous; scutellum extremely small.

Elytral epipleura finely imbricate-punctate; marginal epipleural stria weakly impressed, complete; marginal elytral stria well impressed, slightly carinate, continued as shortened apical elytral stria, next obliterated; humeral elytral stria weakly impressed on basal fourth; inner subhumeral stria well impressed, carinate and almost complete, abbreviated on apical sixth; elytral disc with four thin dorsal elytral striae 1–4, abbreviated basally, apically surpassing half of elytral length, fourth dorsal elytral stria slightly shortened apically, sutural elytral stria absent; elytral disc microscopically punctate, punctation becomes more prominent in postero-lateral corners.

Propygidium almost completely exposed, with lateral depressions, imbricate-punctate, punctures separated by about their own diameter; pygidium likewise imbricate-punctate, punctures becoming sparser medio-apically.

Anterior margin of median portion of prosternum (Fig. 496) rounded, pre-apical foveae absent; prosternal process flat, substrigulate; carinal prosternal striae narrowing anteriorly; lateral prosternal striae carinate, narrowing anteriorly; both sets of striae apically attaining anterior margin of prosternal process.

Anterior margin of mesoventrite deeply emarginate medially; discal marginal mesoventral stria anteriorly carinate, weakened laterally; disc of mesoventrite transverse, smooth; mesometaventral sutural stria absent. Intercoxal disc of metaventrite slightly convex, smooth; lateral metaventral stria weakly impressed, obliquely arcuate, shortened; lateral disc of metaventrite (Fig. 497) concave, imbricate-punctate; metepisternum + fused metepimeron (Fig. 497) likewise imbricate-punctate, but punctation coarser and denser.

Intercoxal disc of the first abdominal sternite laterally depressed, completely striate laterally; imbricate-punctate.

Protibia (Figs. 499–500) slender, flattened, outer margin with sparse row of short thin denticles; setae of outer row extremely short and sparse; setae of median row similarly short and sparse, irregular; protarsal groove absent; anterior protibial stria inconspicuous; two thin short tarsal denticles present apically; protibial spur large, bent, growing out from apical protibial margin; apical margin of protibia posteriorly with two tiny apical denticles; outer part of posterior surface of protibia (Fig. 500) imbricate, with two scattered irregular rows of microscopic sclerotized setae; delimitation from median part of posterior surface vague; posterior protibial stria complete, with rather dense sclerotized setae that become thicker towards apical margin; inner margin with double row of short setae.

Mesotibia (Figs. 160–161) slender, outer margin with a single row of sparse thin denticles growing in size apically; setae of outer row extremely short and minute; setae of median row inconspicuous; posterior mesotibial stria inconspicuous; anterior surface of mesotibia imbricate, with two rows of well sclerotized short setae; anterior mesotibial stria complete, terminating in single tiny inner anterior denticle; mesotibial spur stout, short; apical margin with two tiny denticles; mesotarsus shorter than mesotibia; claws of apical tarsomere longer than half its length; metatibia (Fig. 498) basically similar to mesotibia, but denticles of outer margin almost absent, only near tarsal insertion two minute denticles present; claws of apical tarsomere about half its length.

Male genitalia. Eighth sternite (Figs. 501–502) not fused medially, vela without setae; eighth tergite and eighth sternite not fused laterally (Fig. 503). Ninth tergite (Figs. 504–505) longitudinally divided medially, tenth tergite (Fig. 504) with two moderately long thin setae (styli?); spiculum gastrale (Fig. 504) expanded on both ends. Aedeagus (Figs. 506–507) slender, conspicuously long, approximately 1.5 times as long as 8 th sternite; basal piece of aedeagus short, ratio of its length: length of parameres 1: 6; parameres fused along their basal three-fourths; aedeagus apically slightly curved ventrad (Fig. 507).