Tybalmia pupillata (Pascoe, 1859)

(Figs 24–35)

Hypselomus pupillatus Pascoe, 1859: 35 .

Hypsioma (Jamesia) bipunctata Jekel, 1861: 260 .

Tybalmia pupillata; Thomson, 1868: 45.

Tybalmia orbis Dillon & Dillon, 1945: 177 . Syn. nov.

Note: see full references on Monné (2023) and Tavakilian & Chevillotte (2022).

Remarks. Pascoe (1859) described Hypselomus pupillatus based on a single specimen from Brazil (Pará). A few years later, Jekel (1861) described Hypsioma (Jamesia) bipunctata based on syntypes from French Guiana. Bates (1865) synonymized these two species names. Dillon & Dillon (1945) described T. orbis based on a single female from Bolivia. According to them: “Most closely related to T. pupillata in coloration and in having two tubercles on each humerus, but the yellow macula of elytra is much enlarged and elongate, at least equal in size to the dark macula; humeri with posterior tubercle less prominent, the anterior one is directed slightly anteriorly; the fourth, sixth, eighth, and tenth antennal segments broadly annulate at base; the pronotal tubercle ends in a long process; the antennal tubercles are distinctly separated at base, and the front is not subtriangular.” However: 1) The yellow pubescent macula on the elytra is very variable in size, from almost absent to proportionally large, including in specimens from Bolivia (Figs 27, 33); 2) the shape and size of the lateral tubercles of the prothorax is variable; in addition, the shape and the size of the humeral tubercles is also variable in the other species of the genus; 3) the whitish pubescent ring on the antennomeres is variable, including in specimens from Bolivia (Figs 27, 33); 4) the size of the lateral tubercles of the humerus is variable and, often, do not differ in specimens (males and females) with small or large yellowish pubescent macula; 5) the distance between antennal tubercles appears to be slightly larger in specimens from Bolivia (Figs 27, 28, 32, 33); however, the distance is also another variable feature as, for example, in specimens from French Guiana (see photographs on Bezark 2023); 6) the shape of the frons is practically identical in all specimens examined and may be identical in specimens from Bolivia and other areas, independently of the size of the yellowish elytral pubescent macula. Therefore, as we could not find a reliable difference between T. orbis and T. pupillata, the former is synonymized with the latter.

Current geographical distribution. Colombia, Ecuador, Peru, Venezuela, Guyana, French Guiana, Brazil (Amazonas, Pará).

Material examined. COLOMBIA, Villavicencio (there are at least three places with this name in three Colombian departments), 1 male, 1920, no collector indicated (MZSP) . ECUADOR, Napo: Pacto Sumaco, 1500 m, 1 male, 2 females, 22. XI.2022, J. Vlasak leg. (JVCO) . PERU: Rio Toro, 1 female, no more data (MZSP) . Junín: Satipo, 1 male, IV.1945, A. Maller leg. (MZSP) . SURINAME (new country record): Nassau Mountains, 1 male, 12. III.1949, D.C.Geijskes leg. (MZSP) . BOLIVIA, El Beni:Uyapi, Guanay , 1male, X-XI.1962, no collector indicated (MZSP) . Cochabamba: Chaparé, Sajta [Valle Sajta] , 1 female, III.1993, no collector indicated (MZSP) . BRAZIL, Amazonas: Benjamin Constant, Rio Javari, 1 male, XI.1960 (MZSP, formerly Diringshofen collection) ; 1 female, II.1961 (MZSP, formerly Diringshofen collection) ; 1 male, XII.1961 (MZSP, formerly Diringshofen collection) ; 1 male, 1 female, IV.1962 (MZSP, formerly Diringshofen collection) ; 1 male, V.1962 (MZSP, formerly Diringshofen collection); S„o Paulo de Olivença, Rio Solimões, 1 female, III.1961 (MZSP, formerly Diringshofen collection) ; 1 male, 1 female, no date indicated (MZSP, formerly Diringshofen collection) ; 1 male, IV.1961 (MZSP, formerly Diringshofen collection); Tefé, 1 male, II.1959 (MZSP, formerly Diringshofen collection) ; 1 male, XI.1962 (MZSP, formerly Diringshofen collection). Pará: Óbidos, 1 male, II.1958 (MZSP, formerly Diringshofen collection) .