Walshina esmaraldensis gen. et sp. nov.

Figures 4.1-4.4, 5.2, 5.5, 6.1-6.19

zoobank.org/ 9E482E98-74DA-4ECC-9BE1-6AA10C07D41F

Phenacolemur cf. shifrae Mason, 1990; Walsh, 1996.

Holotype. LACM 40198, left M 1.

Hypodigm. M 2: SDSNH 42268, 62850, 87336, 87337; M 3: SDSNH 76267; M 1: SDSNH 76337, 87331; M 2: SDSNH 76338, 87332, 87333; M 3: SDSNH 72583, 76266, 76339, 87334, 87335.

Type locality. LACM (CIT) 180, north of Simi Valley, Ventura County, California, USA. Late Uintan (Ui3) .

Etymology. In reference to the localities of Emerald Ridge (Vulgar Latin: esmaraldus).

Diagnosis. Intermediate in size between W. mcgrewi comb. nov. and W. shifrae comb. nov. Differs from W. mcgrewi comb. nov. in having a hypoconulid closer to the hypoconid than to the entoconid on M 2, similar to W. shifrae comb. nov. Very large hypoconulids on M 1 and M 2, whereas they are weakly developed in W. mcgrewi comb. nov. and W. shifrae comb. nov. M 2 trigonid and talonid of similar width, whereas W. mcgrewi comb. nov. and W. shifrae comb. nov. have significantly wider M 2 talonid bases relative to the trigonid base. M 3 with larger paraconid and taller metaconid relative to the protoconid than in W. shifrae comb. nov. Hypoconulid lobe of M 3 not expanded as far distally from the apex of the hypoconulid as in W. shifrae comb. nov. Differs from W. mcgrewi comb. nov. (but not W. shifrae comb. nov.) in having upper molars with smaller distolingual basins. In mesial view, M 1 with a less clearly delineated preprotocrista, like W. mcgrewi comb. nov. M 2 with a much more lingually expanded protocone lobe than in W. shifrae comb. nov. (but not in W. mcgrewi comb. nov.).