tonnoiri species-group

Diagnosis Adults: smaller darker species, female total body length 2.2–2.9 mm; Rs neither thickened nor branched distally; junction of r-m cross vein and R 1 with markedly expressed dark spot; pedisulcus more strongly developed as deeper furrowing of cuticle on dorsal base of hind tarsomere II; tarsal claw basal tooth a third or more length of main talon, claw heel slightly expressed; genital fork with anterior stem markedly broadened and bent; male gonostylus cone-shaped, with two apical spines; ventral plate apex broadly rounded, not markedly directed ventrally; parameres smaller. Pupa: gill with fewer (21–50) filaments; abdominal tuberculation slightly to markedly expressed; pleurites present on segments VI & VII. Larva: four to five hypostomal setae; lateral hypostomal serrations present; posterior tentorial pits small and rounded; prothoracic proleg lateral sclerite V-shaped; anal sclerite with posteroventral arms only; area between posterolateral arms and circlet of hooks with small number of campaniform sensilla; posterior proleg with markedly fewer (1,800 –3,000) hooks; larvae occur in flows of various velocity.