Scaphytopius springerae Arias-Paco & Godoy, sp. nov.

(Figs. 6A–F, 9K, 10J, 11W, 12W, 13W)

Description. Length of male 3.50 mm. General appearance light brown with numerous dark spots widely distributed. Crown 1.4 x longer than basal width between eyes, white with several longitudinal brown spots forming almost continuous band between eyes, and another extending longitudinally to apex (Fig. 6A). Face white, with numerous irregular brown spots scattered widely across gena and frontoclypeus, but fewer spots on lorum and anteclypeus (Fig. 6C). Forewings brown with multiple dark spots widely distributed; around 12 widely distributed irregular white spots (Fig. 6B), with 7–8 veinlets; veins brown, and third apical cell with pale outer edge. Legs light brown with black spots on tibiae and femora.

Male genitalia. Pygofer in lateral view rectangular with rounded apex; brown with irregular pale spots, without macrosetae (Fig. 9K). Subgenital plate triangular, wider at base than apex, much longer than wide, with row of aproximately 5 macrosetae on external margin (Fig. 10J). Connective in ventral view W-shaped. Styles thin and elongated, rectangular-shaped; apophysis short, thin, and rounded at apex; space between preapical lobe and apophysis in L-shape (Fig. 6D). Paraphyses in ventral view long and thin, fused at base, they remain parallel in basal quarter, then diverge, crossing at midlength, and converging again toward apex (Fig. 6D). Aedeagus in lateral view wedged between paraphyses just before they cross (Figs. 6D–E); very short, robust, apically rounded, hammer-shaped with curved handle (Fig. 6E); preatrium inconspicuos; dorsal apodeme longer than shaft and curved toward apex (Fig. 6E); shaft short and robust; in dorsal view tubular (Fig. 6F). Gonopore in apical position.

Diagnosis. Thin and elongated styles in rectangular shape with L-shaped space between apophysis and preapical lobe (Fig. 6D). Aedeagus positioned between paraphyses just before they cross (Figs. 6D–E); shaft very short, tubular and robust, rounded toward apex; dorsal apodeme curved toward apex, longer than shaft, giving hammerlike shape with curved handle (Fig. 6E). This species is distinguished from S. brevis, by the following features: In dorsal view the crown is longer; in lateral view shaft much shorter; dorsal apodeme is 2.4x longer than shaft; apex of shaft rounded (Fig. 6E); junction of dorsal apodeme with shaft rounded, giving triangular appearance (Fig. 6E); in ventral view paraphyses much shorter, crossing at same level as apex of styles.

Distribution. Costa Rica.

Biology. Unknown.

Etymology. Dedicated to Monika Springer, aquatic entomologist renowned for her studies on aquatic insects and freshwater ecosystems in Costa Rica.

Material examined. Holotype. ♂. Costa Rica, Guanacaste, Nicoya, San Antonio, Humedal, Corral de Piedra. 6.VII.2005. INB0004155439 (MNCR)