Aname magnifica sp. nov.

urn:lsid:zoobank.org:act: A00657A1-A048-4FFE-938F-E694CDF15614

Figs 10, 69–70

Diagnosis

Males of A. magnifica sp. nov. can be distinguished from all species for which males are known except A. barrema and A. inimica by a moderate to large body size (carapace length> 4.0 mm), the presence of a copulatory organ with the bulb tapering into a long embolus (embolus length / bulb length>1.5) with a slightly thicker basal section that narrows and curves sharply at about 0.6 of length, a patch of thicker setae proximally of asetose depression, and the absence of a pronounced, sharp heel on metatarsus I (as in pallida -complex species) (Fig. 69A–Q). Males of A. magnifica can be distinguished from those of A. inimica by the presence of a longer embolus (embolus length / bulb length>1.5; cf. ~ 1.2) (Fig. 69L– M; cf. Fig. 67). Males of A. magnifica can be distinguished from those of A. barrema by the presence of a shorter asetose depression on the palp tibia (depression length / tibia length ~ 0.6; cf. ~ 0.7 in A. barrema) and a thicker metatarsus I (metatarsus I length / width ~ 3.5; cf. ~ 4.5 in A. barrema) (Fig. 69K, Q; cf. Fig. 64).

Females of A. magnifica sp. nov. can be distinguished from all species for which females are known except A. barrema and A. distincta by the presence of spermathecae with two vesicles, with relatively long and straight lateral vesicles (lateral vesicle length/ genitalia width>0.25) that don’t curve medially at their ends, and very short, straight medial vesicles (medial vesicle length / lateral vesicle length <0.5) (Fig. 70D, L). Females of A. magnifica can be distinguished from those of A. barrema and A. distincta by the presence of larger posterior sigilla on the sternum (posterior sigilla length /sternum length ~ 0.18; cf. ~ 0.14) and spermathecae with triangular lateral vesicles with wide bases tapering to narrow ends (Fig. 70G–H, L; cf. Figs 65–66).

Etymology

The specific epithet ‘ magnifica ’ is an adjective derived from the Latin ‘ magnificus ’, meaning ‘great, grand’, or ‘magnificent’, in reference to the large and relatively robust nature of this species.

Type material

Holotype

AUSTRALIA – New South Wales • ♂; Inverell; 29°46′ S, 151°07′ E; 10 Oct. 1996; L. Abra leg.; AMS KS49693.

Paratypes

AUSTRALIA – New South Wales • 1 ♀; Inverell; 29°46′ S, 151°07′ E; Sep. 1986; R. Gunning leg.; AMS KS16758 • 1 ♀; Inverell; 29°46′ S, 151°07′ E; 10 Aug. 1953; R.F. Cook leg.; AMS KS69998 • 1 ♂; Inverell; 29°46′ S, 151°03′ E; 10 Oct. 1986; University of New South Wales leg.; QMB S6712 .

Other material examined

AUSTRALIA – New South Wales • 1 ♂; Graman; 29°31′ S, 150°20′ E; 6 Sep. 1979; under eucalypt, in garden; QMB S1282 (paratype of Aname inimica Raven, 1985) • 1 ♀; Gurley; 29°44′ S, 149°48′ E; 5 Oct. 2005; AMS KS92856 .

Description

Male (holotype, AMS KS49693)

GENERAL (Fig. 69A–Q). Body length 22.17, in good condition, colour presumably faded due to preservation.

DORSAL PROSOMA (Fig. 69A, E–F). Carapace length 8.92, width 7.47, length/width 1.19, clypeus to fovea length/carapace length 0.72, caput width/carapace width 0.67, carapace dark red-brown, reflective setae present, moderate on caput, moderate on thorax, fovea procurved, fovea width/carapace length 0.16 (Fig. 69A, F); chelicerae dark red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.47 (Fig. 69A); eye group rectangular, width/length 1.91, eye tubercle present (Fig. 69E).

ABDOMEN (Fig. 69B, D). Abdomen length 8.79, light grey, dorsal pattern absent, with consistent cover of short setae.

VENTRAL PROSOMA (Fig. 69C, G–I). Labium cuspules present, count =1 (Fig. 69H); maxillae heel distinct, cuspules present, count = about 130, extending posteriorly onto heel, extending laterally about 35% of maxillae length (Fig. 69C, I); coxae cuspules absent, thorn-like setae on prolateral face present (Fig. 69C, I); sternum length/width 1.16, central sternum with consistent covering of short setae, row of longer setae around posterior edges (Fig. 69G–H); posterior sigilla circular, central sternum to posterior sigilla length/sternum length 0.25, posterior sigilla length/sternum length 0.16 (Fig. 69G–H); other sigilla small, round and lateral (Fig. 69G–H).

LEG I (Fig. 69N–Q). Leg I red-brown, lighter on distal metatarsus and tarsus, reflective setae on dorsal femur, femur length 6.69, patella length 4.25, tibia length 4.70, metatarsus length 4.62, tarsus length 2.94, total length 23.19, leg I length/carapace length 2.60 (Fig. 69N–O); scopulae on distal metatarsus and tarsus (Fig. 69N–O); spine count Fe D 2, Fe PL 1, Pa PL 2, Ti PL 0, Ti RL 0, Me PL 0, Me RL 0, Ta 0 (Fig. 69N–O); tibia length/width [TIL/TID] 3.09, even width along length, spur present, intermediate triangular/digitiform, knuckle absent, megaspine angled at 30 degrees, length to distal face of spur/ tibia length [TIS/TIL] 0.48, spur height/tibia width [TISH/TID] 0.61, megaspine length/tibia length 0.22 (Fig. 69N–P); metatarsus relatively straight, proximal excavation present, excavation concave with slight heel, heel semi-sharp, excavation length/metatarsus length [MIPEL/MIL] 0.48, metatarsus length/ width [MIL/MID] 3.46 (Fig. 69N–O, Q).

PEDIPALP (Fig. 69J–M). Tibia length 3.50, width 1.50, length/width [PTL/PTD] 2.34, asetose depression present, depression length/palp tibia length [PDL/PTL] 0.60, retrolateral face with patch of long setae proximally of asetose depression, ventral face with two elongate bristle-like setae below depression, prolateral face with patch of spines on distal half, disto-medial spine absent (Fig. 69J–K); patella prolateral face with 2 (both rubbed off) spines (Fig. 69J–K); cymbium with scopulae present distally (Fig. 69J–K); copulatory organ total length 2.05, length/palp tibia length 0.59 (Fig. 69L–M); bulb length/ width 1.00 (Fig. 69L–M); embolus tapering from bulb, attenuate, swollen base tapering before strong curve to sinuous tip, one strong bend, at about 0.6 of length, width at base/bulb width 0.33, embolus length/bulb length 1.75 (Fig. 69L–M).

Female (paratype, AMS KS16758)

GENERAL (Fig. 70A–L). Body length 24.90, in good condition, colour presumably faded due to preservation.

DORSAL PROSOMA (Fig. 70A, E–F). Carapace length 9.52, width 8.20, length/width 1.16, clypeus to fovea length/carapace length 0.73, caput width/carapace width 0.78, carapace orange-brown, reflective setae present, light on caput, light on thorax, fovea procurved, fovea width/carapace length 0.16 (Fig. 70A, F); chelicerae red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.57 (Fig. 70A); eye group rectangular, width/length 1.83, eye tubercle present (Fig. 70E).

ABDOMEN (Fig. 70B, D). Abdomen length 9.78, dark brown, dorsal pattern absent, with consistent cover of short setae.

VENTRAL PROSOMA (Fig. 70C, G–I). Labium cuspules absent (Fig. 70H); maxillae heel distinct, cuspules present, count =about 165, extending posteriorly onto heel, extending laterally about 30% of maxillae length (Fig. 70C, I); coxae cuspules absent, thorn-like setae on prolateral face present (Fig. 70C, I); sternum length/width 1.21, central sternum with consistent covering of moderate setae, row of longer setae around posterior edges (Fig. 70G–H); posterior sigilla ovoid, central sternum to posterior sigilla length/sternum length 0.20, posterior sigilla length/sternum length 0.18 (Fig. 70G–H); other sigilla small, round and lateral (Fig. 70G–H).

LEG I (Fig. 70J–K). Leg I orange-brown, femur length 7.04, patella length 5.00, tibia length 4.66, metatarsus length 4.25, tarsus length 2.80, total length 23.75, leg I length/carapace length 2.50; scopulae on distal metatarsus and tarsus; spine count Fe D 1, Fe PL 1, Pa PL 2, Ti PL 2, Ti RL 4, Me PL 1, Me RL 1, Ta 0; tibia length/width [TIL/TID] 2.63.

GENITALIA (Fig. 70D, L). Epigastric furrow unmodified (Fig. 70D); spermathecae with two vesicles each (Fig. 70L); lateral vesicle relatively straight, length 0.74, lateral vesicle length/genitalia width 0.43, length/width at base 1.64, crown un-demarcated (Fig. 70L); medial vesicle short, relatively straight and projecting ventrally, medial vesicle length/genitalia width 0.14, length/width 1.53, medial vesicle length/lateral vesicle length 0.32 (Fig. 70L).

Distribution and natural history

Aname magnifica sp. nov. occurs in north-eastern New South Wales, in the Nandewar and New England Tablelands bioregions, where it is known from four locations extending from Gurley in the east to Inverell in the west (Fig. 10). The form of burrows constructed by spiders of this species is unknown, but is likely to be similar to that of other barrema -complex species (Fig. 10).