Aname harmoniosa sp. nov.

urn:lsid:zoobank.org:act: 3D87A3A5-4536-40E0-BA01-FAE2A402FE5F

Figs 9, 60

Diagnosis

Males of A. harmoniosa sp. nov. can be distinguished from all species for which males are known except A. calida sp. nov., A. carina, A. cassowariensis sp. nov., A. robertsorum, and A. tropicana sp. nov. by a moderate to large body size (carapace length>4.0 mm), and the presence of a long embolus (embolus length / bulb length>1.5) that is thin, with a sharp bend near its base before a relatively straight distal section (Fig. 60L–M). Males of A. harmoniosa can be distinguished from those of A. calida, A. cassowariensis, and A. tropicana by the presence of a longer embolus (embolus length / bulb length>2) (Fig. 60L–M; cf. Figs 56, 59, 62). Males of A. harmoniosa can be distinguished from those of A. carina by the presence of a more proximally-positioned tibial spur (distance to spur/ tibia length <0.6) (Fig. 60P; cf. Fig. 57). Males of A. harmoniosa can be distinguished from those of A. robertsorum by the presence of a longer proximal excavation on metatarsus I (excavation length /metatarsus length ~0.55; cf. ~ 0.44 in A. robertsorum), and a more sinuous embolus (Fig. 60L–Q; cf. Fig. 61).

Females of A. harmoniosa sp. nov. are unknown.

Etymology

The specific epithet ‘ harmoniosa ’ is a Latin adjective meaning ‘harmonious’ or ‘graceful’, in reference to the gracile morphology of this species.

Type material

Holotype

AUSTRALIA – Queensland • ♂; Hann Tableland; 16°49′ S, 145°11′ E; 950–1000 m a.s.l.; 11–14 Dec. 1995; G.B. Monteith, G. Thompson and D.J. Cook leg.; pitfall trap, rainforest; QMB S40518.

Paratypes

AUSTRALIA – Queensland • 1 ♂; Hann Tableland; 16°49′ S, 145°11′ E; 950–1000 m a.s.l.; 11–14 Dec. 1995; G.B. Monteith, G. Thompson and D.J. Cook leg.; pitfall trap, rainforest; QMB S118369 • 2 ♂♂, 1 juv.; Hann Tableland; 16°49′ S, 145°11′ E; 1000 m a.s.l.; 11–13 Dec. 1995; D.J. Cook leg.; pitfall trap, rainforest; QMB S40534.

Description

Male (holotype, QMB S40518)

GENERAL (Fig. 60A–Q). Body length 19.22, in good condition, colour presumably faded due to preservation.

DORSAL PROSOMA (Fig. 60A, E–F). Carapace length 7.67, width 6.15, length/width 1.25, clypeus to fovea length/carapace length 0.70, caput width/carapace width 0.67, carapace dark red-brown, caput slightly darker than thorax, reflective setae present, moderate on caput, light on thorax, fovea procurved, fovea width/carapace length 0.14 (Fig. 60A, F); chelicerae dark red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.54 (Fig. 60A); eye group rectangular, width/length 1.91, eye tubercle present (Fig. 60E).

ABDOMEN (Fig. 60B, D). Abdomen length 6.92, grey, dorsal pattern absent, with consistent cover of short setae.

VENTRAL PROSOMA (Fig. 60C, G–I). Labium cuspules absent (Fig. 60H); maxillae heel distinct, cuspules present, count =about 75, extending posteriorly onto heel, extending laterally about 30% of maxillae length (Fig. 60C, I); coxae cuspules absent, thorn-like setae on prolateral face present (Fig. 60C, I); sternum length/width 1.18, central sternum with consistent covering of short setae, row of longer setae around posterior edges (Fig. 60G–H); posterior sigilla ovoid, central sternum to posterior sigilla length/ sternum length 0.22, posterior sigilla length/sternum length 0.13 (Fig. 60G–H); other sigilla small, round and lateral (Fig. 60G–H).

LEG I (Fig. 60N–Q). Leg I red-brown, lighter on patella, tibia, distal metatarsus and tarsus, reflective setae on dorsal femur, femur length 5.55, patella length 3.79, tibia length 4.39, metatarsus length 4.48, tarsus length 2.48, total length 20.68, leg I length/carapace length 2.70 (Fig. 60N–O); scopulae on distal metatarsus and tarsus (Fig. 60N–O); spine count Fe D 0, Fe PL 1, Pa PL 2, Ti PL 0, Ti RL 0, Me PL 0, Me RL 0, Ta 0 (Fig. 60N–O); tibia length/width [TIL/TID] 3.83, even width along length, spur present, intermediate triangular/digitiform, knuckle absent, megaspine angled at 27 degrees, length to distal face of spur/tibia length [TIS/TIL] 0.58, spur height/tibia width [TISH/TID] 0.69, megaspine length/ tibia length 0.24 (Fig. 60N–P); metatarsus slightly sinuous, proximal excavation present, excavation concave with inconspicuous heel, heel rounded, excavation length/metatarsus length [MIPEL/MIL] 0.55, metatarsus length/width [MIL/MID] 4.62 (Fig. 60N–O, Q).

PEDIPALP (Fig. 60J–M). Tibia length 3.40, width 1.24, length/width [PTL/PTD] 2.73, asetose depression present, depression length/palp tibia length [PDL/PTL] 0.57, retrolateral face with short, thorn-like setae along retrolateral edge of depression, getting denser proximally, ventral face with two elongate bristle-like setae below depression, prolateral face with two disto-ventral spines, disto-medial spine absent (Fig. 60J–K); patella prolateral face with 2 (both rubbed off) spines (Fig. 60J–K); cymbium with scopulae present distally (Fig. 60J–K); copulatory organ total length 1.84, length/palp tibia length 0.54 (Fig. 60L–M); bulb length/width 0.87 (Fig. 60L–M); embolus tapering from bulb, attenuate, very thin, protruding laterally with strong basal curve, one strong bend, at about 0.4 of length, sinuous tip, width at base/bulb width 0.21, embolus length/bulb length 2.15 (Fig. 60L–M).

Distribution and natural history

Aname harmoniosa sp. nov. occurs in northern Queensland, in the Einasleigh Uplands bioregion, on the Hann Tableland, near the town of Mareeba (Fig. 9). The form of burrows constructed by spiders of this species is unknown, but is likely to be similar to that of other robertsorum -complex species (Fig. 9).