Aname olkola sp. nov.

urn:lsid:zoobank.org:act: 1AEC5765-6983-47DE-A36A-A431190F5414

Figs 14, 116–117

Diagnosis

Males of A. olkola sp. nov. can be distinguished from all species for which males are known by a moderate to large body size (carapace length>4.0 mm), the presence of a copulatory organ with a highly rounded bulb and a short embolus (embolus length / bulb length <1.5) that is strongly curved, and a digitiform tibial spur (Fig. 116L–Q).

Females of A. olkola sp. nov. can be distinguished from all species for which females are known by the presence of spermathecae with two vesicles, with relatively wide, bulbous lateral spermathecae (lateral vesicle length / width ~ 1.7), widening towards rounded ends, and short, straight medial vesicles projecting from a position posterior to, and separated from, lateral vesicles (Fig. 117).

Etymology

The specific epithet ‘ olkola ’ is a noun in apposition, referencing Olkola National Park, where the specimens were collected.

Type material

Holotype

AUSTRALIA – Queensland • ♂; Olkola, Valley of Kimba Plateau; 15°15′ S, 143°38′ E; 129 m a.s.l.; Jul. 2015; R.C. Santana leg.; pitfall trap, moist gully; QMB S22650.

Paratype

AUSTRALIA – Queensland • 9 ♂♂, 1 ♀, 1 juv.; Killarney HS, campsite and airstrip; 15°25′ S, 143°30′ E; 165 m a.s.l.; 13–24 Jul. 2015; R.J. Raven, R.C. Santana and L. Carr leg.; open eucalypt forest; QMB S22010.

Description

Male (holotype, QMB S22650)

GENERAL (Fig. 116A–Q). Body length 17.09, in good condition, colour presumably faded due to preservation.

DORSAL PROSOMA (Fig. 116A, E–F). Carapace length 6.81, width 5.67, length/width 1.20, clypeus to fovea length/carapace length 0.71, caput width/carapace width 0.71, carapace dark red-brown, caput slightly darker than thorax, reflective setae present, heavy on caput, moderate on thorax, fovea procurved, fovea width/carapace length 0.19 (Fig. 116A, F); chelicerae dark red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.41 (Fig. 116A); eye group rectangular, width/length 2.42, eye tubercle present (Fig. 116E).

ABDOMEN (Fig. 116B, D). Abdomen length 7.46, brown, dorsal pattern absent, with some evidence of reflective setae, and consistent cover of short setae.

VENTRAL PROSOMA (Fig. 116C, G–I). Labium cuspules absent (Fig. 116H); maxillae heel distinct, cuspules present, count =about 164, extending posteriorly onto heel, extending laterally about 50% of maxillae length (Fig. 116C, I); coxae cuspules absent, thorn-like setae on prolateral face present (Fig. 116C, I); sternum length/width 1.27, central sternum with consistent covering of moderate setae, row of longer setae around posterior edges, some shorter bristle-like setae around anterior edges (Fig. 116G–H); posterior sigilla semi-elongate, central sternum to posterior sigilla length/sternum length 0.24, posterior sigilla length/sternum length 0.14 (Fig. 116G–H); other sigilla small, round and lateral (Fig. 116G–H).

LEG I (Fig. 116N–Q). Leg I red-brown, lighter on distal metatarsus and tarsus, reflective setae on dorsal femur, femur length 5.32, patella length 3.63, tibia length 3.81, metatarsus length 3.86, tarsus length 2.33, total length 18.95, leg I length/carapace length 2.78 (Fig. 116N–O); scopulae on distal metatarsus and tarsus (Fig. 116N–O); spine count Fe D 2, Fe PL 1, Pa PL 2 (distal rubbed off), Ti PL 0, Ti RL 0, Me PL 0, Me RL 0, Ta 0 (Fig. 116N–O); tibia length/width [TIL/TID] 3.25, even width along length, spur present, digitiform, knuckle present, megaspine angled at 35 degrees, length to distal face of spur/ tibia length [TIS/TIL] 0.61, spur height/tibia width [TISH/TID] 0.70, megaspine length/tibia length 0.23 (Fig. 116N–P); metatarsus relatively straight, proximal excavation present, excavation concave with slight heel, heel semi-sharp, excavation length/metatarsus length [MIPEL/MIL] 0.47, metatarsus length/ width [MIL/MID] 3.83 (Fig. 116N–O, Q).

PEDIPALP (Fig. 116J–M). Tibia length 2.55, width 1.14, length/width [PTL/PTD] 2.24, asetose depression present, depression length/palp tibia length [PDL/PTL] 0.58, retrolateral face with patch of long setae proximally of asetose depression, ventral face with two elongate spine-like setae below depression, prolateral face with two disto-ventral spines, disto-medial spine present (Fig. 116J–K); patella prolateral face with 2 spines (Fig. 116J–K); cymbium with scopulae present distally (Fig. 116J–K); copulatory organ total length 1.21, length/palp tibia length 0.47 (Fig. 116L–M); bulb length/width 1.06 (Fig. 116L– M); embolus slightly reflexed, attenuate, tapering and curving relatively evenly to point, one strong bend, at about 0.2 of length, slight bend before tip, width at base/bulb width 0.20, embolus length/bulb length 0.92 (Fig. 116L–M).

Female (paratype, QMB S22010)

GENERAL (Fig. 117A–L). Body in moderate condition except for abdomen, which is completely destroyed.

DORSAL PROSOMA (Fig. 117A, E–F). Carapace length 7.43, width 6.26, length/width 1.19, clypeus to fovea length/carapace length 0.72, caput width/carapace width 0.81, carapace orange-brown, caput slightly darker than thorax, reflective setae present, moderate on caput, light on thorax, fovea procurved, fovea width/carapace length 0.21 (Fig. 117A, F); chelicerae red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.63 (Fig. 117A); eye group rectangular, width/length 2, eye tubercle present (Fig. 117E).

VENTRAL PROSOMA (Fig. 117C, H). Labium cuspules absent (Fig. 117H); maxillae heel distinct, cuspules present, count =about 216, extending posteriorly onto heel, extending laterally about 55% of maxillae length (Fig. 117C); coxae cuspules absent, thorn-like setae on prolateral face present (Fig. 117C); sternum length/width 1.24, central sternum with consistent covering of moderate setae, row of longer setae around posterior edges, some shorter bristle-like setae around anterior edges (Fig. 117H); posterior sigilla semi-elongate, central sternum to posterior sigilla length/sternum length 0.24, posterior sigilla length/sternum length 0.15 (Fig. 117H); other sigilla small, round and lateral (Fig. 117H).

LEG I (Fig. 117J–K). Leg I red-brown, darker on femur, reflective setae on dorsal femur, femur length 5.57, patella length 3.84, tibia length 3.73, metatarsus length 3.36, tarsus length 2.17, total length 18.67, leg I length/carapace length 2.51; scopulae on distal metatarsus and tarsus; spine count Fe D 2, Fe PL 1, Pa PL 2 (proximal rubbed off), Ti PL 2, Ti RL 4, Me PL 2, Me RL 3, Ta 4; tibia length/width [TIL/ TID] 2.75.

GENITALIA (Fig. 117L). Spermathecae with two vesicles each; lateral vesicle relatively straight, widening towards tip, length 0.88, lateral vesicle length/genitalia width 0.33, length/width at base 1.66, crown un-demarcated; medial vesicle short, relatively straight and projecting postero-ventrally, medial vesicle length/genitalia width 0.18, length/width 2.56, medial vesicle length/lateral vesicle length 0.54.

Distribution and natural history

Aname olkola sp. nov. occurs in far northern Queensland, in the Cape York Peninsula bioregion, where it is known from two locations in and around Olkola National Park, west of the town of Laura (Fig. 14). The form of burrows constructed by spiders of this species is unknown.