Aname camara Raven, 1985

Figs 14, 105–107

Aname camara Raven, 1985: 387, figs 3, 15, 23, 27, 37, 54, 63.

Diagnosis

Males of A. camara can be distinguished from all species for which males are known by a moderate to large body size (carapace length>4.0 mm), the presence of a long embolus (embolus length / bulb length> 1.5) that is very strongly curved, and a small, tight patch of thorn-like setae on the retrolateral palp tibia (Figs 105J–M, 106J–M).

Females of A. camara can be distinguished from all species for which females are known except A. aurensis sp. nov., A. briggsi sp. nov., A. dingo sp. nov., A. eddieorum sp. nov., A. longitheca, A. mulgana sp. nov., and A. rupicola sp. nov. by the presence of spermathecae with two vesicles, with relatively long and straight lateral vesicles (lateral vesicle length/genitalia width>0.25) and long medial vesicles (medial vesicle length/ lateral vesicle length>1) that project medially or posteromedially, before undulating towards the anterior (Fig. 107L). Females of A. camara can be distinguished from those of A. aurensis, A. briggsi, A. dingo, A. eddieorum, A. longitheca, A. mulgana, and A. rupicola by the presence of a light tan body colouration and spermathecae with relatively short vesicles (medial vesicle length /genitalia width ~0.3; cf.>0.35) with widely-spaced crowns (distance between crowns is roughly equal to the length of the lateral vesicles) (Fig. 107A–L; cf. Figs 35, 37–38, 40, 43, 45, 47).

Type material

Holotype

AUSTRALIA – Queensland • ♂; Aramac; 22°58′ S, 145°15′ E; 21 Aug. 1960; F. McKenzie leg.; QMB S1250.

Allotype

AUSTRALIA – Queensland • 1 ♀; Aramac; 22°58′ S, 145°15′ E; 21 Aug. 1960; F. McKenzie leg.; QMB S1251.

Other material examined

AUSTRALIA – Queensland • 2 ♀♀; Aramac; 22°58′ S, 145°15′ E; 21 Aug. 1960; F. McKenzie leg.; QMB S1252 • 1 ♂; Bellen Park Station; 23°58′ S, 143°52′ E; 200-250 m a.s.l.; 14 Mar. 1992; A. Emmott leg.; grey cracking clay, mitchell grass downs; QMB S20402 .

Description

Male (holotype, QMB S1250)

GENERAL (Fig. 105A–Q). Body length 17.93, in poor condition, colour dramatically faded, embolus broken, tissue degraded.

DORSAL PROSOMA (Fig. 105A, E–F). Carapace length 6.34, width 5.31, length/width 1.20, clypeus to fovea length/carapace length 0.67, caput width/carapace width 0.63, carapace pallid red-brown, caput much darker than thorax, reflective setae absent or inconspicuous, fovea procurved, fovea width/carapace length 0.13 (Fig. 105A, F); chelicerae orange-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.57 (Fig. 105A); eye group rectangular, width/length 1.92, eye tubercle present (Fig. 105E).

ABDOMEN (Fig. 105B, D). Abdomen length 7.48, light tan, dorsal pattern absent, with consistent cover of short setae.

VENTRAL PROSOMA (Fig. 105C, G–I). Labium cuspules absent (Fig. 105H); maxillae heel distinct, cuspules present, count =about 50, extending posteriorly onto heel, extending laterally about 35% of maxillae length (Fig. 105C, I); coxae cuspules absent, thorn-like setae on prolateral face present (Fig. 105C, I); sternum length/width 1.10, most setae rubbed off, row of longer setae around posterior edges, some shorter thorn-like setae around anterior edges (Fig. 105G–H).

LEG I (Fig. 105N–Q). Leg I pallid, translucent, femur length 5.11, patella length 3.12, tibia length 3.71, metatarsus length 3.41, tarsus length 2.29, total length 17.65, leg I length/carapace length 2.78 (Fig. 105N–O); scopulae on distal metatarsus and tarsus (Fig. 105N–O); spine count Fe D 1, Fe PL 1, Pa PL 4, Ti PL 0, Ti RL 0, Me PL 0, Me RL 0, Ta 0 (Fig. 105N–O); tibia length/width [TIL/TID] 2.73, widening from proximal end to spur before narrowing again towards distal end, spur present, triangular, knuckle absent, megaspine angled at 25 degrees, length to distal face of spur/tibia length [TIS/TIL] 0.42, spur height/tibia width [TISH/TID] 0.47, megaspine length/tibia length 0.25 (Fig. 105N–P); metatarsus slightly sinuous, proximal excavation present, excavation concave with pronounced heel, heel sharp, excavation length/metatarsus length [MIPEL/MIL] 0.57, metatarsus length/width [MIL/MID] 3.09 (Fig. 105N–O, Q).

PEDIPALP (Fig. 105J–M). Tibia length 2.46, width 1.17, length/width [PTL/PTD] 2.10, asetose depression present, depression length/palp tibia length [PDL/PTL] 0.54, retrolateral face with distinct patch of short thorn-like setae proximally, ventral face with two elongate bristle-like setae below depression, prolateral face with two disto-ventral spines, disto-medial spine present (Fig. 105J–K); patella prolateral face with three (one rubbed off) spines (Fig. 105J–K); cymbium with scopulae present distally (Fig. 105J–K); copulatory organ total length 1.44, length/palp tibia length 0.58 (Fig. 105L–M); bulb length/width 0.80 (Fig. 105L–M); embolus demarcated and roughly perpendicular to bulb, attenuate, embolus protruding relatively perpendicular to bulb, with sharp bend just after base, one strong bend, at about 0.3 of length, width at base/bulb width 0.25, embolus length/bulb length 1.91 (Fig. 105L–M).

Male (QMB S20402)

GENERAL (Fig. 106A–Q). Body length 15.83, in good condition, colour presumably faded due to preservation.

DORSAL PROSOMA (Fig. 106A, E–F). Carapace length 6.82, width 5.57, length/width 1.22, clypeus to fovea length/carapace length 0.68, caput width/carapace width 0.63, carapace red-brown, caput much darker than thorax, reflective setae present, moderate on caput, moderate on thorax, fovea procurved, fovea width/carapace length 0.13 (Fig. 106A, F); chelicerae red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.37 (Fig. 106A); eye group rectangular, width/length 1.98, eye tubercle present (Fig. 106E).

ABDOMEN (Fig. 106B, D). Abdomen length 6.46, tan, with grey dorsal strip, dorsal pattern absent, with consistent cover of short setae.

VENTRAL PROSOMA (Fig. 106C, G–I). Labium cuspules absent (Fig. 106H); maxillae heel distinct, cuspules present, count =about 70, extending posteriorly onto heel, extending laterally about 35% of maxillae length (Fig. 106C, I); coxae cuspules absent, thorn-like setae on prolateral face present (Fig. 106C, I); sternum length/width 1.16, central sternum with consistent covering of short setae, row of longer setae around posterior edges, some shorter thorn-like setae around anterior edges (Fig. 106G–H); posterior sigilla semi-elongate, central sternum to posterior sigilla length/sternum length 0.21, posterior sigilla length/sternum length 0.19 (Fig. 106G–H); other sigilla small, round and lateral (Fig. 106G–H).

LEG I (Fig. 106N–Q). Leg I pallid, darker on proximal patella, metatarsus and tarsus, femur length 5.24, patella length 3.24, tibia length 3.75, metatarsus length 3.48, tarsus length 2.22, total length 17.93, leg I length/carapace length 2.63 (Fig. 106N–O); scopulae on distal metatarsus and tarsus (Fig. 106N–O); spine count Fe D 1, Fe PL 1, Pa PL 1, Ti PL 0, Ti RL 0, Me PL 0, Me RL 0, Ta 0 (Fig. 106N–O); tibia length/width [TIL/TID] 2.78, even width along length, spur present, triangular, knuckle absent, megaspine angled at 28 degrees, length to distal face of spur/tibia length [TIS/TIL] 0.43, spur height/ tibia width [TISH/TID] 0.61, megaspine length/tibia length 0.23 (Fig. 106N–P); metatarsus slightly sinuous, proximal excavation present, excavation concave with pronounced heel, heel sharp, excavation length/metatarsus length [MIPEL/MIL] 0.60, metatarsus length/width [MIL/MID] 3.27 (Fig. 106N–O, Q).

PEDIPALP (Fig. 106J–M). Tibia length 2.49, width 1.19, length/width [PTL/PTD] 2.09, asetose depression present, depression length/palp tibia length [PDL/PTL] 0.56, retrolateral face with distinct patch of short thorn-like setae proximally, ventral face with one elongate bristle-like seta below depression, prolateral face with two disto-ventral spines, disto-medial spine present (Fig. 106J–K); patella prolateral face with 3 spines (Fig. 106J–K); cymbium with scopulae present distally (Fig. 106J–K); copulatory organ total length 1.01, length/palp tibia length 0.41 (Fig. 106L–M); bulb length/width 0.78 (Fig. 106L–M); embolus demarcated and roughly perpendicular to bulb, attenuate, embolus protruding relatively perpendicular to bulb, with sharp bend just after base, one strong bend, at about 0.3 of length, width at base/bulb width 0.25, embolus length/bulb length 1.81 (Fig. 106L–M).

Female (allotype, QMB S1252)

GENERAL (Fig. 107A–L). Body length 23.43, in moderate condition, colour faded significantly due to preservation.

DORSAL PROSOMA (Fig. 107A, E–F). Carapace length 6.52, width 5.18, length/width 1.26, clypeus to fovea length/carapace length 0.67, caput width/carapace width 0.65, carapace orange, reflective setae present, light on caput, light on thorax, fovea procurved, fovea width/carapace length 0.14 (Fig. 107A, F); chelicerae red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.63 (Fig. 107A); eye group rectangular, width/length 1.78, eye tubercle present (Fig. 107E).

ABDOMEN (Fig. 107B, D). Abdomen length 11.96, light tan-brown, dorsal pattern absent, with consistent cover of short setae.

VENTRAL PROSOMA (Fig. 107C, G–I). Labium cuspules absent (Fig. 107H); maxillae heel distinct, cuspules present, count =about 54, extending posteriorly onto heel, extending laterally about 50% of maxillae length (Fig. 107C, I); coxae cuspules absent, thorn-like setae on prolateral face present (Fig. 107C, I); sternum length/width 1.15, central sternum with consistent covering of short setae, row of longer setae around posterior edges (Fig. 107G–H); posterior sigilla semi-elongate, central sternum to posterior sigilla length/sternum length 0.23, posterior sigilla length/sternum length 0.17 (Fig. 107G–H); other sigilla small, round and lateral (Fig. 107G–H).

LEG I (Fig. 107J–K). Leg I orange, femur length 4.93, patella length 3.24, tibia length 3.23, metatarsus length 2.82, tarsus length 1.94, total length 16.16, leg I length/carapace length 2.48; scopulae on distal metatarsus and tarsus; spine count Fe D 1, Fe PL 1, Pa PL 2, Ti PL 1, Ti RL 4 (2 proximal weak), Me PL 2, Me RL 3, Ta 0; tibia length/width [TIL/TID] 2.66.

GENITALIA (Fig. 107D, L). Epigastric furrow unmodified (Fig. 107D); spermathecae with two vesicles each (Fig. 107L); lateral vesicle relatively straight, length 0.62, lateral vesicle length/genitalia width 0.29, length/width at base 1.32, crown un-demarcated (Fig. 107L); medial vesicle with distinct basal section angled medially, before undulating towards anterior, medial vesicle length/genitalia width 0.40, length/width 8.3, medial vesicle length/lateral vesicle length 1.37 (Fig. 107L).

Distribution and natural history

Aname camara occurs in central Queensland, in the Mitchell Grass Downs bioregion, roughly between the towns of Stonehenge and Aramac (Fig. 14). The form of burrows constructed by spiders of this species is unknown.

Remarks

A male specimen from further west within the same bioregion (QMB S20402) is here linked with this species. The embolus shape is distinct; however, both emboli of the holotype male are damaged and may not reflect the original form. In all other respects, the two specimens are virtually identical, and we have therefore designated them as conspecific.