Uca (Tubuca) coarctata (H. Milne Edwards, 1852)

(Fig. 5F–H)

Gelasimus coarctatus H. Milne Edwards, 1852: 146 [type locality: “Odessa”]; A. Milne-Edwards 1873: 272.

Uca dubia — Sakai 1974: 10, 13; Sakai 1976b: 38.

Uca (Deltuca) coarctata coarctata — Crane 1975: 52, figs. 9B, 26A, 31A, 36B, 37C, 38M–P, 62A–C, 81D, 82H, pl. 6E–H, 7, 46C–D; Sakai 1976a: 603, text-figs. 328, 330, pl. 208(2); Sakai 1976b: 32, 38, frontispiece fig. 2; Shih 1994: 73, figs. 48– 50.

Uca coarctata — Fukui et al. 1989: 227, fig. 2; Huang et al. 1989: 194, fig. 3, pl. 2A–D; Yamaguchi 1994: 159; Wang & Liu 1996a: 46, 2 unnumbered figs.; Lee 2001: 101, 5 unnumbered figs.; Ng et al. 2001: 37 [list]; Yoshigou 2001: 4, fig. 2, pl. 1E; Liu & Wang 2010: 33, 2 unnumbered figs.; Japanese Association of Benthology 2012: 214, 1 unnumbered fig.; Fujita & Uyeno 2015: 99, fig. 5A–C.

Uca coarctata coarctata — Yamaguchi 1994: 183; Nakasone & Irei 2003: 269, fig. 29C, D.

Uca dussumieri — Katsu 2007: 82, 4 unnumbered figs (not Gelasimus dussumieri H. Milne Edwards, 1852).

Uca crassipes — Katsu 2007:90, 1 unnumbered fig.

Uca (Tubuca) coarctata —Ng et al. 2008: 241 [list]; Toyota & Seki 2014: 229, 4 unnumbered figs.

Material examined. Taiwan: 1 ♀ (16.8 mm) (ASIZ), Lanyang R. estuary, Yilan, coll. H.-C. Liu, 25 July 2004 ; 2 ♂♂ (12.7, 18.6 mm) (NCHUZOOL 14679), Yanshuei R. estuary, Tainan, 4 Aug. 2009; 1 ♀ (18.4 mm) (NCHUZOOL 14685), Yanshuei R. estuary, Tainan, 18 Oct. 2011; 1 ♂ (17.5 mm), 1 ♀ (21.7 mm) (TMCD 2026), Baoli R. estuary, Pingtung, 26 Apr. 1983; 1 ♂ (18.4 mm) (TMCD CHCD 780), Baoli R. estuary, Pingtung, coll. H.- C. Liu, 20 Apr. 1995; 2 ♂♂ (13.5, 14.5 mm) , 3 ♀♀ (12.7–15.0 mm) (TMCD CHCD 810), 4 ♂♂ (12.1–19.2 mm), 2 ♀♀ (8.8, 13.3 mm) (TMCD CHCD 812), 6 ♂♂ (12.0– 24.4 mm), 1 ♀ (14.2 mm) (TMCD CHCD 813), Baoli R. estuary, Pingtung, 14 June 1995; 1 ♂ (22.4 mm) (NCHUZOOL 14683), 1 ♀ (20.0 mm) (NCHUZOOL 14681), Baoli R. estuary, Pingtung, coll. H.- T. Shih, 12 Jan. 2000; 2 ♂♂ (13.2, 13.2 mm) (NCHUZOOL 14682), Baoli R. estuary, Pingtung, coll. H.-T. Shih, 20 July 2011. Penghu Islands: 1 ♀ (21.0 mm) (NCHUZOOL 13231), Citou, coll. H.- T. Shih & Y.-H. Wang, 19 May 2007 .

Distribution. Western Pacific (Ryukyus, Taiwan [including Penghu], China (?), Philippines, Indonesia [including Sumatra], New Guinea, Australia).

Remarks. Crane (1975) examined specimens of U. coarctata from Luzon, Philippines, which was the northernmost record in the western Pacific in her monograph. Sakai (1976a, b), however, identified his “ U. dubia ” (Sakai 1974) as U. coarctata, although Gelasimus dubius Stimpson, 1858, had been confirmed by Crane (1975) as U. dussumieri (see below). The two species can be distinguished by the morphology of orbital floor and major fingers: U. coartctata has the orbital floor with a row of tubercles (without tubercles in U. dussumieri) and the dactylus of the major cheliped with 1 groove (with two grooves in adult U. dussumieri).

The first records of this species in Taiwan are those of Fukui et al. (1989) and Huang et al. (1989). Shih et al. (2008) confirmed it from Penghu. Katsu (2007) identified a female from Amami, central Ryukyus, as “ U. dussumieri ”, but it is U. coarctata because there is a row of tubercles on the floor of the orbit.