Subtribe Bolitocharina Thomson, 1859
0445. Bolitochara Mannerheim, 1830: 75
Type species: Aleochara pulchra Gravenhorst, 1806
Agaribiota Bierig, 1937b: 279 [synonymized by Blackwelder 1952: 41]
Type species: Bolitochara cinctigastra Bierig, 1937
Ditropalia Casey, 1906: 263 [synonymized by Seevers 1978: 164]
Type species: Bolitochara bella Märkel, 1844
3 species are included in this genus from the West Indies and South America.
Characters/Description: Fenyes 1920: 114 (characters as Ditropalia); Coiffait & Sáiz 1967: 65 (characters in key), 79 (characters); Ashe 1992: 365 (characters in key), 367 (characters).
Key: Coiffait & Sáiz 1967: 80 (Chilean species).
Distribution.
West Indies: Cuba.
South America: Colombia, Argentina.
Other regions: Palaearctic, Oriental, Australian, Oceanic.
0446. Habramicrada Pace, 2007: 225 (Fig. 58)
Type species: Habramicrada boliviensis Pace, 2007
4 species are known from and are restricted to South America.
Distribution.
South America: Peru, Bolivia, Brazil.
0447. Leptusa Kraatz, 1856b: 60
Type species: Bolitochara pulchella Mannerheim, 1830
Subgenera:
Adexiopisalia Pace, 1982c: 585
Type species: Leptusa hummleri Bernhauer, 1910
Adoxopisalia Pace, 1989: 130
Type species: Leptusa opaca Casey, 1893
Afroleptusa Pace, 1983c: 62
Type species: Leptusa sparsa Fauvel, 1898
Akratopisalia Pace, 1996a: 29
Type species: Leptusa cribrata Pace, 1996
Amalopisalia Pace, 1983c: 97
Type species: Leptusa janetscheki Scheerpeltz, 1956
Anisopisalia Pace, 1989: 214
Type species: Leptusa luigionii Bernhauer, 1935
Anomopisalia Pace, 1983c: 82
Type species: Leptusa granulicauda Eppelsheim, 1890
Anosiopisalia Pace, 1995: 247
Type species: Leptusa nemoricultrix Pace, 1995
Aphaireleptusa Pace, 1996a: 30
Type species: Leptusa anmashanensis Pace, 1996
Aleteleptusa Pace, 1997a: 753 [Smetana 2004: 429]
Type species: Leptusa chinensis Pace, 1997
Mimumenepisalia Pace, 1997a: 752 [Smetana 2004: 429]
Type species: Leptusa gonggamontis Pace, 1997
Baldopisalia Pace, 1983e: 130
Type species: Leptusa veronensis Pace, 1983
Boreoleptusa Pace, 1989: 82
Type species: Leptusa norvegica Strand, 1941
Bothrydiopisalia Scheerpeltz, 1966b: 20
Type species: Leptusa bosnica Eppelsheim, 1892
Bryopisalia Pace, 1989: 239
Type species: Leptusa pilatensis Scheerpeltz, 1972
Chondrelytropisalia Scheerpeltz, 1976c: 55
Type species: Leptusa nepalica Scheerpeltz, 1976
Chondridiopisalia Pace, 1983c: 85
Type species: Leptusa granulifera Pace, 1983
Chondridiopisalia Scheerpeltz, 1966b: 19 [nomen nudum, Newton & Thayer 2003]
Type species: Leptusa granulifera Scheerpeltz, 1966
Chondridiopisalia Lohse, 1974: 53 [Newton & Thayer 2003]
Chondropisalia Scheerpeltz, 1966b: 14
Type species: Leptusa schaschli Ganglbauer, 1897
Cidnopisalia Pace, 1989: 235
Type species: Leptusa cavallensis Pace, 1980
Comphopisalia Scheerpeltz, 1966b: 16
Type species: Leptusa brucki Scriba, 1867
Craspedopisalia Scheerpeltz, 1966b: 19
Type species: Leptusa carinata Bernhauer, 1909
Cryptopisalia Pace, 1983c: 88
Type species: Leptusa montisparii Scheerpeltz, 1953
Cryptopisalia Pace, 1983e: 129 [Newton & Thayer 2003]
Cyllopisalia Pace, 1982a: 40
Type species: Leptusa difformis Mulsant & Rey, 1853
Cyllopisalia Pace, 1983c: 79 [Smetana 2004: 431]
Type species: Leptusa difformis Mulsant & Rey, 1853
Parapisalia Scheerpeltz, 1966b: 18 [synonymized by Pace 1982a: 40]
Type species: Leptusa difformis Mulsant & Rey, 1853
Sipalia Mulsant & Rey, 1853b: 32 [synonymized by Pace 1982a: 40]
Type species: Homalota difformis Mulsant & Rey, 1853
Dendroleptusa Pace, 1983c: 65
Type species: Leptusa fumida Erichson, 1839
Drepanoleptusa Pace, 1982d: 93
Type species: Leptusa annapurnensis Pace, 1982
Dysleptusa Pace, 1982c: 580
Type species: Oxypoda fuliginosa Aubé, 1850
Ectinopisalia Scheerpeltz, 1966b: 18
Type species: Leptusa baldensis Ganglbauer, 1895
Emmelopisalia Pace, 1983c: 92
Type species: Leptusa norica Pace, 1979
Entomophallopisalia Pace, 1983c: 90
Type species: Leptusa asturiensis Eppelsheim, 1880
Eospisalia Pace, 1982c: 583
Type species: Leptusa ishizuchiensis Pace, 1982
Eremopisalia Pace, 1989: 196
Type species: Leptusa marianii Pace, 1978
Eucryptusa Casey, 1906: 345
Type species: Silusa nanula Casey, 1893
Evaniopisalia Pace, 1982a: 38
Type species: Leptusa ultracollensis Pace, 1982
Exaeretopisalia Pace, 1989: 233
Type species: Leptusa cansigliensis Pace, 1975
Gallopisalia Pace, 1989: 179
Type species: Leptusa lapidicola Brisout de Barneville, 1863
Gatineauleptusa Klimaszewski in Klimaszewski et al., 2018: 363
Type species: Leptusa gatineauensis Klimaszewski & Pelletier in Klimaszewski et al., 2004
Geopisalia Pace, 1984: 214
Type species: Leptusa peyerimhoffi Bernhauer, 1935
Gnopheropisalia Pace, 1983c: 76
Type species: Leptusa granulipennis Eppelsheim, 1880
Habropisalia Pace, 1989: 216
Type species: Leptusa laticeps Scheerpeltz, 1972
Halmaeusa Kiesenwetter in Kiesenwetter & Kirsch, 1877: 160 [*]
Type species: Halmaeusa antarctica Kiesenwetter, 1877
Antarctophytosus Enderlein, 1909: 377 [synonymized by Steel 1964: 366]
Type species: Phytosus atriceps Waterhouse, 1875
Paraphytosus Cameron, 1917c: 125 [synonymized by Steel 1964: 366]
Type species: Phytosus atriceps Waterhouse, 1875
Austromalota Brèthes, 1925: 170 [synonymized by Steel 1964: 366]
Type species: Austromalota rufimixta Brèthes, 1925
Hemipasilia Pace, 1989: 90
Type species: Leptusa oertzeni Eppelsheim, 1888
Heteroleptusa Pace, 1989: 160
Type species: Sipalia frontalis Casey, 1893
Heterotyphlopasilia Pace, 1983c: 66
Type species: Leptusa serbica Scheerpeltz, 1935
Homopisalia Pace, 1982c: 584
Type species: Leptusa bengalensis Pace, 1982
Ischnopisalia Pace, 1989: 226
Type species: Leptusa poggii Pace, 1978
Kochliodepisalia Pace, 1996a: 35
Type species: Leptusa spirarum Pace, 1996
Koilokoryphepisalia Pace, 1999a: 210
Type species: Leptusa argentierensis Pace, 1999
Lasiopisalia Pace, 1983c: 76
Type species: Leptusa crenulata Bernhauer, 1899
Lathripisalia Pace, 1983e: 135
Type species: Leptusa montispasubii Pace, 1975
Leptopasilia Scheerpeltz, 1966b: 13
Type species: Leptusa hopffgarteni Eppelsheim, 1881
Leptusa Kraatz, 1856b: 60
Type species: Bolitochara pulchella Mannerheim, 1830
Sipalia Mulsant & Rey, 1853a: 45 [Newton & Thayer 2003]
Liopisalia Pace, 1989: 177
Type species: Leptusa serullazi Sainte-Claire Deville, 1919
Megacolypisalia Pace, 1983e: 133
Type species: Leptusa laevicauda Scheerpeltz, 1958
Melopisalia Pace, 1983c: 88
Type species: Leptusa pasubiana Scheerpeltz, 1953
Metaplasmopasilia Pace, 1989: 104
Type species: Leptusa apennina Holdhaus, 1924
Microcolypisalia Pace, 1989: 196
Type species: Leptusa occulta Pace, 1975
Micropisalia Scheerpeltz, 1948: 164
Type species: Leptusa alpestris Scheerpeltz, 1935
Myopopisalia Scheerpeltz, 1966b: 14
Type species: Leptusa bonvouloiri Brisout de Barneville, 1867
Nanoglossa Fauvel, 1868a: 379 [*]
Type species: Microglossa chilensis Fauvel, 1866
Microglossa Fauvel, 1866b: 282 [synonymized by Fauvel 1868a: 379]
Type species: Microglossa chilensis Fauvel, 1866
Microglossa Fauvel, 1866a: 33 [Newton & Thayer 2003]
Type species: Microglossa chilensis Fauvel, 1866
Pareuryusa Scheerpeltz, 1972d: 131 [synonymized by Pace 1989: 110]
Type species: Euryusa parallela Fairmaire & Germain, 1862
Nanopisalia Pace, 1983c: 94
Type species: Homalota piceata Mulsant & Rey, 1853
Micropisalia Scheerpeltz, 1966b: 20 [synonymized by Pace 1983c: 94]
Type species: Homalota piceata Mulsant & Rey, 1853
Nanopisalia Pace, 1983e: 132 [Newton & Thayer 2003]
Type species: Homalota piceata Mulsant & Rey, 1853
Neopisalia Scheerpeltz, 1966b: 16
Type species: Leptusa microphthalma Reitter, 1887
Nesopisalia Pace, 1992b: 112
Type species: Leptusa centralis Pace, 1992
Stenoleptusa Scheerpeltz, 1966b: 10 [synonymized by Assing 2009c: 1286]
Type species: Bolitochara laeviuscula Hochhuth, 1849
Oligopisalia Scheerpeltz, 1966b: 20
Type species: Leptusa flavicornis Brancsik, 1874
Oncopisalia Pace, 1982c: 589
Type species: Leptusa monachorum Bernhauer, 1912
Oreopisalia Pace, 1983c: 82
Type species: Leptusa hoelzeli Scheerpeltz, 1948
Pachygastropisalia Scheerpeltz, 1966b: 15
Type species: Leptusa lativentris Sharp, 1873
Pachygluta Thomson, 1858: 34
Type species: Oxypoda ruficollis Erichson, 1839
Parapisalia Scheerpeltz, 1948: 159
Type species: Leptusa puellaris Hampe, 1863
Pasilia Mulsant & Rey, 1871: 226
Type species: Homalota nubigena Kiesenwetter, 1861
Pasilia Mulsant & Rey, 1872b: 316 [Newton & Thayer 2003]
Pasilia Mulsant & Rey, 1873c: 73 [Newton & Thayer 2003]
Pisalia Mulsant & Rey, 1871: 234
Type species: Homalota globulicollis Mulsant & Rey, 1853
Pisalia Mulsant & Rey, 1872b: 324 [Newton & Thayer 2003]
Pisalia Mulsant & Rey, 1873c: 73 [Newton & Thayer 2003]
Tropidiopasilia Scheerpeltz, 1966b: 11 [synonymized by Pace 1989: 190]
Type species: Leptusa tricolor Scriba, 1870
Planepisalia Pace, 1989: 217
Type species: Leptusa kochiana Bernhauer, 1936
Protoleptusa Semenov in Semenov et al., 2001: 156
Type species: Leptusa defuncta Semenov, Perkovsky & Petrenko, 2001
Rhombopisalia Scheerpeltz, 1953: 66
Type species: Leptusa knabli Bernhauer, 1909
Roubaliusa Scheerpeltz, 1966b: 13
Type species: Leptusa storkani Roubal, 1916
Scelopisalia Scheerpeltz, 1966b: 18
Type species: Leptusa gracilipes Krauss, 1899
Scoliophallopisalia Pace, 1989: 232
Type species: Leptusa pascuorum Pace, 1975
Scoliophallopisalia Pace, 1981: 115 [nomen nudum, Newton & Thayer 2003]
Scoliophallopisalia Anonymous, 1984: 249 [Newton & Thayer 2003]
Type species: Leptusa blesioi Pace, 1981
Scolioplatypisalia Pace, 1989: 86
Type species: Leptusa munelensis Apfelbeck, 1907
Stictopisalia Scheerpeltz, 1966b: 16
Type species: Leptusa eximia Kraatz, 1856
Synpisalia Scheerpeltz, 1966b: 16 [synonymized by Pace 1983c: 56]
Type species: Leptusa caucasica Eppelsheim, 1877
Toxophallopisalia Pace, 1983c: 99
Type species: Leptusa austriaca Scheerpeltz, 1935
Trichopasilia Scheerpeltz, 1966b: 12
Type species: Leptusa major Bernhauer, 1900
Typhlopasilia Ganglbauer, 1895: 278
Type species: Leptusa kaufmanni Ganglbauer, 1895
Ulitusa Casey, 1906: 347
Type species: Ulitusa cribratula Casey, 1906
Yunnaleptusa Assing, 2008b: 256
Type species: Leptusa curvata Assing, 2006
16 species are included in this genus from South America.
Characters/Description: Lynch-Arribálzaga 1884: 87 (characters); Fenyes 1920: 120 (characters), 122 (characters subgenera Typhlosilia and Leptusa), 123 (characters subgenera Eucryptusa and Pachygluta), 124 (characters subgenera Oreusa and Pasilia), 125 (characters subgenera Pisalia); Pace 1989: 25 (characters), 110 (characters of the subgenus Nanoglossa), 244 (characters of the subgenus Halmaeusa).
Key: Bernhauer 1900a: 402 (Subgenera and some species); Fenyes 1920: 121 (subgenera); Pace 1989: 25 (subgenera), 110 (species of the subgenus Nanoglossa), 246 (species of the subgenus Halmaeusa); Pace 2001a: 12 (Chilean species).
Distribution.
South America: Chile, Argentina, Falkland Isl.
Other regions: Nearctic, Palaearctic, Ethiopian, Oriental, Australian, Antarctic.
0448. Meoticaphaena Pace, 2015e: 72
Type species: Meotica cordobensis Bernhauer, 1934
Only one species is known from and is restricted to South America.
Distribution.
South America: Argentina.
0449. Phymatura J. Sahlberg, 1876: 85
Type species: Bolitochara brevicollis Kraatz, 1856
Venusa Casey, 1906: 272 [synonymized by Ashe 1992: 362]
Type species: Venusa picta Casey, 1906
3 species are included in this genus from South America.
Characters/Description: Fenyes 1920: 116 (characters); Ashe 1992: 366 (characters in key), 371 (characters). Distribution.
South America: Brazil, Argentina.
Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian.
0450. Prosoponotha Pace, 1999c: 146
Type species: Prosoponotha chilensis Pace, 1999
5 species are known from and are restricted to South America.
Distribution.
South America: Brazil, Chile.
0451. Tanystetha Pace, 2002: 137
Type species: Tanystetha brasiliensis Pace, 2002
Only one species is known from and is restricted to South America.
Distribution.
South America: Brazil. Subtribe Dinardopsina Bernhauer & Scheerpeltz 1926
0452. Dinardopsis Bruch, 1917: 257 (Fig. 52)
Type species: Dinardopsis solenopsidicola Bruch, 1917
2 species are known from and are restricted to South America.
Distribution.
South America: Brazil, Argentina.