Glyptapanteles arcanus species group

The large G. arcanus species group forms two separate clades in the current phylogeny (Fig. 2) and is therefore clearly not a monophyletic group. There is one small clade of two species ( G. ruhri Fagan- Jeffries, Bird & Austin sp. nov. and G. rodriguezae Fagan-Jeffries, Bird & Austin sp. nov. plus an undescribed lineage, with a sequence in BOLD) which is sister to the G. niveus species group clade, whilst the remainder of species are paraphyletic with respect to the G. mouldsi species group and include G. arcanus Fagan-Jeffries, Bird & Austin sp. nov., G. doreyi Fagan-Jeffries, Bird & Austin sp. nov., G. erucadesolator Fagan-Jeffries, Bird & Austin sp. nov., G. goodwinnoakes Fagan-Jeffries, Bird & Austin sp. nov., G. lambkinae Fagan-Jeffries, Bird & Austin sp. nov., G. lessardi Fagan-Jeffries, Bird & Austin sp. nov., G. vergrandiacus Fagan-Jeffries, Bird & Austin sp. nov. and G. wrightae Fagan-Jeffries, Bird & Austin sp. nov. The species group, as currently recognised, is distributed throughout the eastern coast of Australia, from the northern regions of QLD through to central NSW and the ACT. One species, G. erucadesolator sp. nov., is also known from northern NT (Fig. 4B).

Species in the G. arcanus species group can be separated from the other described species of Glyptapanteles in Australia by the combination of the following characters: gena without a pale spot, T1 and T2 dark, anteromesoscutum punctures not extremely sparse and deep (i.e., not as in the niveus species group), no orange markings on anteromesoscutum postero-laterally, propodeum either strongly sculptured (rugose) or with punctures in the anterior half and shallow rugose sculpturing in the posterior half, ventral side of antennal scape (at least in distal half) paler than flagellomeres, tegula pale.