Propsilocerus paradoxus (Lundström)

(Figs. 40–48)

Orthocladius paradoxus Lundstrȍm, 1915: 14 .

Synpropsilocerus paradoxus Kieffer 1923: 3 .

Syndiamesa komensis Zvereva, 1950: 275; Pankratova 1970: 75.

Propsilocerus paradoxus (Lundstr̂m); Saether & Wang 1996: 469; Petrova et al. 2003: 36; Saether 2004: 17; Makarchenko & Makarchenko 2006: 349; Ashe & O’Connor 2012: 502.

Material examined. Russia: 2 adult males, Eastern Siberia, Trans-Baikal Territory, Chita City, Kenon Lake, 1.V.2003, 52.030147 N, 113.380136 E, leg. О. Klishko ; 4 adult males, Jewish Autonomous Region, Smidovichsky District, Nature Reserve “Bastak”, Zabelovskoye Lake, 48.432583 N, 134.213889 E, 16.IV.2008, leg. V. Lubarets ; 2 adult males, Republic of Buryatia, Selenginsky District, Gusinoe Lake, 20.V.2017, leg. N. Bazova.

Comments. Some variation is added based on the males from the Russian Far East.

Description

Adult male (n = 4, except when otherwise stated). Total length 4.2–6.2 mm (n = 11). Total length/wing length 1.24–1.36.

Coloration (slide-mounted material). Head, thorax, tergites of abdomen dark brown; legs brown ot yellowishbrown; hypopygium brown; wings grayish.

Head. Eyes bare, reniform. Temporal setae including 2–3 verticals and without postorbitals. Clypeus with ca 50–70 setae. Antenna with 13 flagellomeres and developed plume; terminal flagellomere with 3 sensitive setae, 20–36 μm long; AR 2.25–2.33. Palpomeres lengths (in μm): 20–40; 68–73; 152–160; 92–100; 100–108. Palpomeres 1–5 length/head width 1.68–1.74.

Thorax. Antepronotum with 13–21 lateral setae. Acrostichals absent, dorsocentrals 60–84 (in anterior part in 2–3 rows, in posterior in 4–6 rows), prealars 6–11 scutellars 21 – 28 (in 1–2 rows);

Wing. Length 3.4–4.1 mm, width 0.84–1.2 mm. R with 8 setae, R 1 and R 4+5 with 0 setae. Costa extension 120–190 μm long. RM length/MCu length 1.22–1.3. Anal lobe developed, outline rounded. Squama with 27 setae 72–92 μm long. VR 1.0–1.1.

Legs. Spur of fore tibia 84–96 μm long; spurs of mid tibia 56–60 μm and 64–72 μm; of hind tibia 56–62 μm and 72–92 μm long. Lengths and proportions of leg segments as in Table 7.

Hypopygium (Figs. 40–48). Tergite IX with 102–122 setae and tapering to pointed apex anal point 68–132 μm long; with 5–7 lateral setae, 36–48 μm long (Figs. 46–48). Laterosternite IX with 9–12 setae. Phallapodeme 129–180 μm long; transverse stemapodeme 208–264 μm long, slightly concave, with oral projections (Fig. 43). Gonocoxite 590–804 μm long. Inferior volsella as in Figs. 40–41), 400–432 μm long. Median volsella with 2–7 strong setae 64–100 μm long on small tubercle (Figs. 42, 45). Gonostylus with two branches, basal branch 188–246 μm long; megaseta 8 μm long (Figs. 40, 44). HR 3.28 –4.9.

Remarks. Comparison of males from the Russian Far East with males from other populations showed that they are similar in main characters, except for a greater number of dorsocentrals (60–84), a longer anal point (68–132 μm) and a transverse sternapodeme (208–264 μm). Specimens from China have 36–73 dorsocentrals, the length of anal point is 40–79 µm, and length of transverse sternapodeme is 120–184 µm (Table 8).

Distribution. Reliably known from China, Mongolia, East Siberia and Russian Far East.