Quemedice enigmaticus Mello-Leitão, 1942

Figs. 1–12, 17–28

Quemedice enigmaticus Mello Leitão, 1942: 407, figs 31–32 (female holotype, examined). Roewer, 1954: 790 (sub. Thomisidae); Platnick, 2008 (sub. Philodromidae).

Type material: Holotype: female from Puna [27°48' S; 62°31' W], Santiago del Estero Province, Argentina, M. Birabén (MLP 15450).

Other material examined. BRAZIL. Mato Grosso: 1ɗ, Chapada dos Guimarães, Usina Hidrelétrica de Rio Manso, Rio Manso [14°52'00" S; 55°48'00" W], 2000–2001, Equipe Rio Manso (IBSP 28197). Minas Gerais: 1Ψ, Montes Claros [16°43'00" S; 43°52'00" W], 2005, G.L.D. Leite (IBSP 73206); 1ɗ, Juatuba, Rio Manso, Serra Azul, Área de Proteção de Mananciais da COPASA [20°10’00" S; 44°21’00" W], 18–24 April 2002, Equipe Biota (IBSP 67823); 1ɗ (IBSP 67822), 2Ψ (IBSP 67821); 1Ψ (IBSP 67820). São Paulo: Castilho 1Ψ, [20°52'00" S; 51°28'60" W], 27 November 1964, Exp. CDZ (MZSP 3898); 1Ψ, Assis [22°40'00" S; 50°25'00" W], Nov.1964, G. Brizola (IBSP 1899); 1ɗ, Itupeva [23°08'60" S; 47°04'00" W], October 1982, I. Levis (IBSP 4160); 1Ψ, São Paulo [23°31'60" S; 46°37'00" W], October1952, J.J. Rufino (IBSP 824); ARGENTINA. Santiago del Estero: 1ɗ, Parque Nacional Copo [25°55'00" S; 61°55'00" W], 22–24 February 2004, C. Grismado, A. Ojanguren, F. Labarque and L. Compagnucci (MACN-Ar 10758).

Diagnosis. Quemedice enigmaticus is distinguished from Q. piracuruca n. sp. by the male palp with RTA branches similar in size (sometimes dorsal branch slightly shorter) and base of embolus smooth (Figs 8–10); female epigynum with broad median septum, twice longer than wide, and vulva with large, elongate hyaline receptacles and slender, convoluted copulatory ducts (Figs 11–12).

Description. Male (IBSP 67822): Prosoma pale orange with brown margins, a median longitudinal brown stripe and a pair of slightly lighter paramedian ones (Figs 5–7). Chelicerae pale orange with small brown spots at the base of setae. Legs orange, except femora with dorsal longitudinal brown stripes (Figs 5– 7). Pedipalps pale orange. Labium pale brown (Fig. 7). Endites and sternum yellow with pale brown margins (Fig. 7). Opisthosoma cream colored with three longitudinal brown stripes (Figs 5–7). Total length 11.1. Dorsal shield of prosoma: 3.8 long, 2.5 wide. Eye diameters and interdistances: AME 0.22, ALE 0.18, PME 0.16, PLE 0.16, AME–AME 0.20, AME–ALE 0.04, PME–PME 0.20, PME–PLE 0.20, AME–PME 0.22, ALE– PLE 0.24. Legs (2413): I: femur 6.9, patella 1.5, tibia 6.7, metatarsus 6.8, tarsus 2.1, total 24.0; II: 8.40, 1.60, 8.10, 8.40, 2.30, 28.80; III: 4.00, 1.30, 3.50, 3.30, 1.10, 13.20; IV: 8.10, 1.40, 6.70, 7.70, 1.90, 25.80. Spination as in genus description, except tibiae III d0–0–1; metatarsus IV d1–0–0.Opisthosoma: 7.5 long, 1.5 wide. Epiandrous spigots dispersed along the margin of epigastric furrow (Fig. 28).

Redescription. Female (holotype, MLP 15450): Coloration as in male (Figs 1–2, 4). Total length 9.62. Dorsal shield of prosoma: 3.96 long, 2.68 wide. Eye diameters and interdistances: AME 0.20, ALE 0.16, PME 0.16, PLE 0.14, AME–AME 0.24, AME–ALE 0.10, PME–PME 0.24, PME–PLE 0.32, AME–PME 0.26, ALE–PLE 0.26. Legs (2413): I: femur 5.50, patella 1.54, tibia 4.32, metatarsus 4.64, tarsus 2.12, total 18.12; II: 7.35, 1.96, 5.74, 5.66, 2.12, 22.83; III: 3.16, 1.12, 2.68, 2.20, 1.00, 10.16; IV: 8.65, 1.48, 4.77, 5.25, 1.40, 21.55. Spination: as in genus description except femur I d0-1-0; r0-1-1; femur IV r0-1-1.Opisthosoma: 5.66 long, 2.88 wide.

Variation. Three males: total length 11.0–11.1; dorsal shield of prosoma 3.7–3.8; femur I 6.9–7.3. Three females: total length 13.1–16.4; carapace 3.8–4.1; femur I 5.7–6.7.

Distribution. Central eastern South America: Argentina and Brazil (state of Mato Grosso).