Eadya annleckieae Ridenbaugh sp. n. Figs 2A-C; 3A-E

Diagnosis.

Eadya annleckieae sp. n. can be distinguished from all other members of Eadya by the following combination of characters: Clypeus flanged across ventral margin, without medial tubercles (Fig. 3A); frons with weak inter-antennal carinae and lateral carina with a faint elevated ridge wrapping around the antennal socket (Fig. 3A, B); occipital carina simple (Fig. 3B); occiput normal; notaulus wide and rugulose (Fig. 3C); scutellar sulcus divided into two distinct foveae with rugulose sculpturing along the posterior margins (Fig. 3C); sternaulus rugulose (Fig. 3D); propodeum rounded in appearance from lateral angle, without transverse carinae (Fig. 3E), and not creating a distinct posterior face when viewed laterally; propodeal spiracle circular; head black except for mandible orange with base black and apex ferruginous, maxillary and labial palp orange (Figs 2A; 3A), antenna dark brown (Figs 3C); pronotum black (Figs 2B; 3B); propleuron black (Fig. 3D); hindwing hyaline with dark brown veins (Fig. 2C); legs orange except for hind tibia dark orange with apex black (Fig. 2A); amino acid sequence (112-118) LRRLTNI (Fig. 15).

Description.

Female. Body length 6.46mm. Ovipositor length 1.72mm.

Color. Head black except for mandible orange with base black and apex ferruginous, maxillary and labial palp orange, and antenna dark brown (Figs 2A; 3A, C); prothorax black (Fig. 2A); mesoscutum black (Fig. 2B); mesopleuron black with the dorsal posterior margin orange (Fig. 3D); scutellum black except for the posterior margin directly behind the scutellar sulcus orange (Fig. 2B); sternum black; metathorax orange (Fig. 2A); forewing and hindwing hyaline with dark brown veins (Fig. 2C); legs orange except for hind tibia dark orange with apex black (Fig. 2A, B); abdomen orange except for ovipositor sheath brown (Fig. 2A, B).

Head. Clypeus simple, punctate and pubescent, flanged across ventral margin, without medial tubercles (Fig. 3A); mandibles overlapping, dorsal and ventral teeth of equal length (Fig. 3A); face densely punctate, pubescent (Fig. 3A); frons rugulose, with a weak inter-antennal carinae and with lateral carinae with a faint elevated ridge wrapping around the antennal socket (Fig. 3A, B); vertex punctate and pubescent (Fig. 3B); occipital carina simple (Fig. 3B), reaching the hypostomal carina; hypostomal carina simple, not strongly flanged, meeting the mandible at the mandibular condyle; occiput smooth, normal.

Mesosoma . Pronotum exposed in dorsal view, pronope and subpronobe present, covered in rugulose sculpturing (Fig. 3C, D); mesoscutum with posterior half of median mesonotal lobe rugulose, a distinct longitudinal carinae extending from the posterior margin to about the middle of the lobe (Figs 2B; 3C); notaulus wide and rugulose (Figs 2B; 3C); scutellar sulcus divided into two distinct foveae with rugulose sculpturing along the posterior margins (Figs 2B; 3E); sternaulus rugulose (Fig. 3D); propodeum rugose, covered in setae but not pubescent, rounded in appearance from lateral angle, without transverse carina and not creating a distinct posterior face when viewed laterally (Figs 2A; 3D, E); propodeal spiracle circular; coxa, trochanter, trochantellus, and femur covered in setae, tibia and tarsus pubescent (Fig. 2A, B); tarsal claws simple.

Forewing. r-m sinuous (Fig. 2C).

Hindwing. R1a with three hamuli.

Metasoma. Metasomal tergite 1 petiolate, spiracle protruding as a tubercle at about the middle of the segment, dorsal surface smooth, lateral surface punctate with associated setae; ovipositor straight (Fig. 2A).

Male. Same as female.

Host.

Paropsisterna nobilitata (Erichson, 1842), Paropsisterna variicollis *, Paropsisterna selmani Reid & de Little, 2013, Paropsis charybdis .

Variations.

Paratype with propleuron black except for lateral posterior margin orange; mesoscutum orange except for the median mesonotal lobe black with the anterior margin and lateral mesonotal lobes ferrunginous (Fig. 3C); mesopleuron orange except for the sternaulus and ventral margins black; scutellum orange (Fig. 3C, E); legs orange except for apex of hind tibia black and hind tarsus with tarsomere 1 yellow and white at apex, tarsomeres 2-4 white, and tarsomere 5 yellow; abdomen orange except for lateral margins of metasomal sternites 3-6 brown, the second and third to last metasomal tergites with two light brown spots near the anterior margin. Some of this variation may be the result of the DNA extraction process.

Diagnostic molecular characters.

Amino acid positions (22-27) MWAGIL; (32-34) SII; (41-46) SRGSLL; (54) R; (67-73) MVMPVIM; (81) I; (90) I; (95-98) MNNM; (104-109) LPSLFI; (112-118) LRRLTNI; (126) I; (133-139) GGRHSGV; (143-144) VA; (150) I; (157) [I or K]; (167-169) FNM; (172-191) NGIAVDRVTLFRWSVKITAF (Fig. 13).

Distribution.

Tasmania.

Etymology.

This species is named in honor of the science fiction author, Ann Leckie by the second author (EB).

Remarks.

This species is referred to as Eadya sp.1 in Peixoto et al. (2018). The UCFC paratype is in poor shape due to the DNA extraction process. The flange of the inter-antennal carinae is difficult to see in the images (Fig. 3A, B), but is clear when viewing the specimens, provided the antennae are separated enough.

Type material.

Holotype, Female (ANIC), "Ellendale, TAS, Female, 21a, 10 Dec 2014, D. Satchell". Paratype, Female (ANIC), "Moina, TAS, 41°29.5'S, 152°04.7'E, Paropsis charybdis sentinel, Emerged 2 Jan 2013, G.R. Allen, E127", "DNA voucher BJS196", GenBank accession numbers KX031361, KX99032, and KX990052. Paratype, Male (UCFC), "The Lea, TAS, #12, Eadya paropsidis cocoon (brown). Emerged from Pst. variicollis *, 4 Dec 2014, UCFC 0 567 827", "DNA voucher BJS501", GenBank accession number KX990216.