B. grandis Lawrence, 1965
Fig. 5
Bicoxidens grandis Lawrence, 1965; Krabbe, 1982; Hamer, 1998; Mwabvu, 2000, 2005 Rhodesiostreptus matabele Schubart, 1966 syn. n.
Bicoxidens matabele Krabbe, 1982; Hamer, 1998; Mwabvu, 2000
Type material B. grandis (examined): Holotype: ZIMBABWE: 1 ɗ, 15 km West of Birchenough Bridge [19°50'S, 32°30'E], 20.ii.1962, D. G. Broadley (NMSA 8884)
Type material B. matabele (examined): Holotype: ZIMBABWE: 1 ɗ, WN West Nicholson, 6.v.1951 (MZLU /L951/3730)
Additional material examined: ZIMBABWE: 1 ɗ, Silobela [19º20' S, 29º35'E], xi.1997, T. Mwabvu (NMZ uncatalogued); 1 ɗ 2Ψ, Birchenough Bridge [19°55'S, 32°35'E], 1998, T. Mwabvu (NMSA 20528); 1 ɗ 3Ψ, Zvomukonde [20º50'S, 30º30'E], xii.1998, T. Mwabvu (NMZ uncatalogued); 2 ɗ 4Ψ, Silobela [19º20' S, 29º30'E], xii.1998, T. Mwabvu (NMSA 20546).
Diagnosis: Apical telocoxite with a horizontal apically acute lateral process and a curved median process that overlaps with median process from opposite gonopod (Figs. 5 a, 5b). Aborally, an apical ear-shaped lobe present on lateral leaf at same level as axe-shaped median process.
Description:
Size: Body length 135 mm; minimum and maximum body width 7–8 mm and 8–11 mm; antenna length 7–9 mm
Number of body rings: 52–62.
Colour: Body black; antennae and legs brown.
Collum: With three complete folds (Fig. 5 d).
Pre-femoral process of 1st pair of male legs: Proximal halves parallel, almost touching; rounded distally, with apical nipples deflected to face the other (Fig. 5 c).
Sigilla on body ring: Three rows, becoming one ventrally.
Gonopod: 6–7 mm long. Orally, apical telocoxite with a lateral and median process (Fig. 5 a). Median process crescent-shaped and overlapping the opposite process. Lateral process horizontal, long and apically acute. Distally, median leaf with median axe-shaped process not touching the opposite one (Fig.5 a).Femoral lobe typically long, (Fig. 5 a).
Aborally, a small almost ear- shaped transverse projection is present on the lateral leaf (Fig. 5b) at same level as the axe-shaped process (Fig. 5b).
Distribution: Current records are from southern Zimbabwe (Fig.10)
Remarks: The gonopod of Rhodesiostreptus matabele Schubart, 1966 is identical to that of B. grandis, and the taxa also have overlapping distributions. Additionally, they both have an ear-shaped projection on the lateral leaf behind the median leaf apex, as well as identical telocoxites and median leaves and a telopodite with a much longer femoral lobe unlike the other species of Bicoxidens . B. matabele is therefore a junior synonym of B. grandis .
Although grandis is larger than flavicollis, their gonopods are similar in that they have overlapping, flattened median processes on the distal telocoxites and axe-shaped processes on the median leaf unlike the other species. The two species are, however, quite distinct; flavicollis does not have a lateral process on the apical telocoxite or an aboral ear-shaped projection on the lateral leaf. Unlike flavicollis, the paracoxite of grandis is not tightly attached to the lateral leaf as in friendi and gokwensis . B. grandis has only been recorded in hot and dry areas.
Abbreviations: asp, axe-shaped process; at, apical telocoxite; cf, collum fold; fl, femoral lobe; llp, lateral leaf projection; ll, lateral leaf; lp, lateral process; mp, median process; pc, paracoxite; pfl, prefemoral lobe; st, sternite; tp, telopodite