Incertae sedis in the Formicinae

Genus † Kyromyrma . Comparative morphological study of † Kyromyrma (∼92 Ma, New Jersey amber; Grimaldi & Agosti, 2000) at the gross (Figs 8L, 9K) and fine scales reveals considerable morphological affinity to Lasius (holotype examined at AMNH). In the original description of † Kyromyrma, the authors did not address the problem of within-subfamily placement, merely noting that ‘the fossil bears an overall resemblance to Prolasius, mostly by virtue of the generalized morphology’ (Grimaldi & Agosti, 2000, p. 13681). Our combined evidence analyses resulted in ambiguous support for the placement of † Kyromyrma, with the genus being recovered as sister to the Lasius genus group (Figure S5), sister to the core Lasiini (Figures S8, S 9), sister to all Lasiini (Figure S7), or sister to Formicinae exclusive of Myrmelachistini (Fig. 4). Statistical support for these placements was uniformly low.