Chlaenius (Sphodromimus) enleensis Mandl, 1992 stat. nov.

Figs 8, 10A, B, 16, 24, 32A-F, 33A-E, 40

Chlaenius flavofemoratus enleensis Mandl, 1992: 100; Lorenz 1998: 318 (synonymized with C. flavofemoratus Laporte,1834, catalogue); Lorenz 2005: 338 (catalogue); Kirschenhofer 2017: 491 (as a subspecies of C. flavofemoratus; catalogue); Azadbakhsh and Kirschenhofer 2019: 1 (transferred to subgenus Chlaenius Sphodromimus from subgenus Chlaenius Haplochlaenius).

Chlaenius tamdaoensis Kirschenhofer, 2003: 32 (type locality: Vietnam, Tam Dao; genus Chlaenius, subgenus Chlaenius Haplochlaenius); Lorenz 2005: 342 (catalogue); Azadbakhsh and Kirschenhofer 2019: 1 (genus Chlaenius, subgenus Chlaenius Sphodromimus) syn. nov.

Type locality.

Indo Chine.

Material examined.

Total 5 specimens. Vietnam: Holotype, Male (NHMB), Indo Chine coll. Dussault/ En-Le 1908/ Chl ( Chlaenius Macrochlaeniles) Chlaenius Macrochlaeniles flavofemoratus ssp. enleensis Dr. K. MANDL det. 1978/ Holotype [red label]. Paratype (DWC, photo) , 1 ♂ (IZAS), Vietnam, Tam Dao, 20-28.VI.1990, Dr. Blazicek lgt./ Paratypus, Chlaenius (Haplochlaenius) tamdaoensis mihi det. Kirschenhofer 2001[red label]/COLL WRASE, BERLIN ; 1 ♂ (IZAS), Vietnam, Tam Dao, 60 km NW Hanoi, 900 m, 1997 May-June, S. Ryabov [internal sac fully everted] ; 1 ♂ (IZAS), Vietnam, Cao Bang, Nguen Binh, 800 m, 2003.V.13, S. Ryabov [internal sac partially everted] ; 1 ♂ (IZAS), Tonkin, Hoa-Binh, leg. A. de Cooman [genitalia damaged by dermestes beetle] .

Diagnosis.

Pronotum green to coppery. PW/PL = 1.06-1.12; PAW/PBW = 0.82-0.95 (Fig. 24); pronotum cordate with anterior angles rounded, not projected forward; disc sparsely punctate and pubescent, with shallow transverse rugosities. Elytral intervals gently convex throughout; densely punctate and pubescent. Hind wings reduced. Metepisterna short; MW/ML = 1.11-1.17 (Fig. 16). Apex of femora dark brown or yellow-brown, the rest of legs black.

Description.

BL = 21.7-24.1 mm, BW = 8.2-8.7 mm. PL = 4.5-4.9 mm, PW = 5.0-5.3 mm, MW = 2.0-2.1 mm, ML = 1.7-1.8 mm. Head, elytra, venter dark and black; pronotum green to coppery; antennae, labial and maxillary palpi, apex of mouthparts and tarsomeres dark brown; apex of femora dark brown or yellow-brown, the rest of legs black.

Head. Vertex finely punctate, pubescent, without a distinct glabrous area; antennae long, reaching middle of elytra; antennomere 3 ~ 1.5 × longer than antennomere 4.

Pronotum cordiform, PW/PL = 1.06-1.12 (Fig. 24), widest at apical third; anterior margin slightly concave, PAW/PBW = 0.82-0.95; lateral margins distinctly narrowed from middle to base, slightly sinuate before posterior angles; anterior angles rounded, not projected forward; posterior angles nearly right angled, rounded at tips; disc gently convex, sparsely punctate and pubescent, with shallow, transverse rugosities, without glabrous area; median line distinct, fine, not reaching anterior margin and base; basal foveae deeply arcuate, punctate and pubescent.

Elytra elongate, EL/BW = 1.58-2.22, gently convex near anterior third, widest near posterior third, rounded at apex in males; striae with deep punctures; parascutellar striae well developed; parascutellar pores present; intervals gently convex throughout, densely punctate and pubescent; sutural angles sharp at tips; hind wings reduced.

Venter densely punctate, pubescent, metepisterna (Fig. 16) short, MW/ML = 1.11-1.17; abdominal sternites III-VI with one setiferous puncture each side, sternite VII with one pair of setiferous punctures in males; all abdominal sternites with distinct impressions laterally.

Legs long and slender; tarsi nearly smooth dorsally.

Male genitalia. Median lobe (Figs 32B-E, 33B-E) long, strongly bent to ventral side; apical orifice opened dorsally, long and wide, not reaching basal bulb; in dorsal view, apical lamella linear, longer than basal width (Figs 32A, 33A), distinctly bent to right side, with a denticulation laterally in the middle, left denticulation distinctly larger than the right one; in left lateral view, apical portion slightly bent dorsally at apex, basal orifice ~ 90° relative to preapical shaft; left paramere larger than right paramere, both helically curved; endophallus (Fig. 32F) with flagellum helically thick and straight; basal part of flagellum with irregular bursa; apical part of flagellum with triangular sclerite.

Female genitalia unknown.

Distribution.

(Fig. 40) Vietnam (Indo Chine). We mark En-le, Yunnan on the map with a question mark ‘?’ .

Remarks.

Lorenz (1998) and Lorenz (2005) proposed C. flavofemoratus enleensis Mandl, 1992 as a synonym of C. flavofemoratus, but Kirschenhofer (2017) treated it as a distinct subspecies. Based on its original description, its pronotum longer than C. flavofemoratus, and black femora indicated it probably represented a different species. After the examination of the holotype and its dissected genitalia, we find that it has the same apical lamella as C. tamdaoensis . As a consequence of this surprising discovery, the locality should also be critically revised. The labels of the holotype contain two localities ( “En-le” and "Indo Chine"), the latter including today’s Vietnam. Based on the type locality of C. tamdaoensis, it is very unlikely that C. flavofemoratus enleensis also occurs in Yunnan’s En-le. Hence, we think that the label of En-le is likely to be the wrong one and may have been erroneously added, as it rarely happened in the historical collections of the NHMB (e.g., Caldara et al. 2022). We upgrade C. enleensis as a valid species and consequently treat C. tamdaoensis as synonym of C. enleensis .