Drypetes madagascariensis (Lam.) Humbert & Leandri

Bull. Mus. Hist. Nat. (Paris), sér. 2, 4: 119 (1932). — Ilex madagascariensis Lam., Encycl., III: 148 (1789).

— Type: Commerson s.n. (holo-, P).

Cyclostemon aquifolium Scott-Elliot, J. Linn. Soc., Bot. 29: 49 (1891). — Drypetes aquifolium (Scott-Elliot) Pax & K. Hoffm., in Engl., Pflanzenr. 81: 277 (1922).

— Type: Scott-Elliot 2873 (holo-, P).

Drypetes madagascariensis subvar. inermis Humbert & Leandri, Bull. Mus. Hist. Nat. (Paris), sér. 2, 4: 119 (1932).

— Type: Humblot 551 (holo-, P; iso-, MO).

Drypetes madagascariensis var. perrieri Humbert & Leandri, Bull. Mus. Hist. Nat. (Paris), sér. 2, 4: 119 (1932).

— Type: Perrier de la Bâthie 18260 (lecto-, P, designated here).

DISTRIBUTION. — Widespread in Madagascar.

With highly variable leaves (their margins often spiny-dentate), but distinguished by its glabrous petioles, 5 sepals, 3(-4) stamens, and 2 carpels.