Scymbalium anale (NORDMANN 1837)

(Fig. 1)

Material examined: France: 1ex., " Grimont " (SDEI) ; 6 exs., inundation of Gers river (MHNG, SMTD) ; 1 ex., inundation of Gers river, XII.1866 (SMTD) ; 2 ex., Gard, leg. Puel (MHNG) ; 5 exs., locality not specified (SDEI, SMF, SMNS); 1 ex., " Gallia merid. " (SMTD). French or Spanish territory: 1 ex., " Pyrenées " (cBor). Austria: 2 exs., Burgenland, Neusiedlersee (SMF). Italy: Lombardia: 3 exs., Rocca de Giorgi (PV), 14.IV.2000, leg. Monzini (ZMUC). Emilia - Romagna: 2 exs., Ferrara, IV.1970, leg. Bordoni (cBor) ; 1 ex., Argenta (FE), Oasi di Campotto, light trap, 4.-5.VI.2005, leg Fabbri (cBor). Toscana: 2 exs., Firenze, Sesto Fiorentino, III.1912, leg. Boglione (MCSNT) ; 1 ex., Grosseto, Poggio Cavallo [42°46'N, 11°10'E], III.1906, leg. Moncini (MCSNM) ; 1 ex., Pistoia, NE Quarrata, La Querciola Caserana [ca. 43°51'N, 11°00'E], 27.IV.2001, leg. Bordoni (cBor). Lazio: 1 ex., Maccanese, leg. Luigioni (SMF). Basilicata: 2 ex., Policoro (MT), 25.I.1976, leg. Angelini (SMNS) ; 4 exs., Matera, Bosco Policoro, I.1976, leg. Montemurro (cBor, cAss) ; 2 exs., Matera, Gravina F. Bradano, light trap, 26.VII.1993, leg. Angelini (ZMUC, cAss). Calabria: 5 exs., Antonimina, 1905, leg. Paganetti (SMF, SMNS, SMTD) ; 1 ex., Brindisi, leg. Leder, Reitter (SMTD) ; 1 ex., Gerace, leg. Paganetti (SMTD). Puglia: 2 exs., Brindisi, leg. Reitter & Leder (SMF). Sardegna: 2 exs., Paulilatino, II.1943, leg. Falzoni (MCSNT) ; 2 exs., Sassari, Chilivani, 25.IV.1979, leg. Pavesi (MCSNM) ; 2 exs., Cagliari, Guspini, VI.1987, leg. Meloni (cBor) ; 1 ex., locality not specified (SMF). Sicilia: 1ex., BoscodiFicuzza (PA), 6.II.1970, leg. Aliquó (cBor). Locality not specified: 1ex.," Italia "(HNHM. Slovakia: 5 exs., Kamen. most, 3.V.1955, leg. Smetana (cBoh, cAss). Romania: 1 ex., " Comana Vlasca ", leg. Montandon (SMTD) ; 1 ex., locality not specified (SMTD). Croatia: 1 ex., Istria, " Val d. Quieto ", 6.IV.1925 (SMF). Bulgaria: 1 ex., Pomorie env., 9.-18.V.1985, leg Wrase (SMNS) ; 1 ex., Promorie env., 9.V.1985, leg. Jaeger (cAss). Albania: 13 exs., Kolonjë, 3 km ENE Leskovik, 40°10'N, 20°37'E, 990 m, flooded calcareous pasture with stones, under stones, 28.V.2010, leg. Assing (cAss). Greece: Mainland 8 exs., ca. 20 km SSE Lamia, Oros Kallidromo, 38°44'N, 22°34'E, 1100m, partly flooded meadow, under stones, 2.IV.2001, leg. Assing (cAss) ; 24 exs., same data, but 6.IV.2001 (cAss); 28 exs., Oros Kallidromo, 38°45'N, 22°30'E, 990m, flooded meadow, under stones, 7.IV.2001, leg. Assing (cAss) ; 2 exs., Parnassos Oros, near Delfi, 1200 m, 11.IV.1989, leg. Karner (cAss). Lesbos: 1 ex., Megali Limni, 39°06'N, 26°20'E, meadows, under stones, 24.III.2005, leg. Lompe & Meybohm (cAss). Turkey: Istanbul: 1ex.,Istanbul (MHNG). Kocaeli / Sakaria: 1ex., Goek Da÷Õ, leg. Bodemeyer (SMF). Bursa: 1 ex., Karacabey, leg. Ajtai (NHMW). Kastamonu: 1ex.,ca. 65kmWKastamonu, 20 km W Eflani, 41°28'N, 33°13'E, 1090 m, calcareous arable land, under stones, 25.III.2010, leg. Assing (cAss). Sinop: 1ex.,15 kmSWSinop, S KÕlÕçlÕ, 41°57'N, 35°02'E, 80 m, grassy road margin, grass between shrubs, sifted, 3.IV.2009, leg. Assing (cAss). Çank Õ r Õ: 2 exs., pass between ÇankÕrÕ and Ilgaz, 1400 m, 3.V.1987, leg. Giachino (cAss, cVai). Manisa: 1 ex., Hamidiye, 8.IV.2007, leg. Anlaú (cAss) ; 1 ex., Hamidiye, 8.IV.2007 (cAss). Ankara 277 exs., 57.5 km SE Bolu, ca. 20 km N BeypazarÕ, 40°18'N, 31°59'E, 1540 m, wet fallow near shallow pond, under stones, 28.III.2010, leg. Assing (cAss, cFel, cSch, MNHUB) ; 1 ex., Ankara, S Ankara, 3 km S Bezirhane, 24.IV.1993, leg Slaven (cPüt). Mu ÷ l a: 1 ex., 100 km W Antalya, 30 km SSE Gölhisar, 1600 m, meadow, 22.V.1993, leg. Schulz (cAss). Isparta: 4 exs., 10 km SE Sütçüler, 37°25'N, 31°02'E, 1520 m, moist meadow, under stones, 26.IV.2011, leg. Assing (cAss) ; 2 exs., 43 km ESE E÷irdir, Dedegöl Da÷larÕ, 5 km W Yeniúarbademli, 37°43'N, 31°19'E, 1820 m, flooded pasture and calcareous slope, under stones, 25.IV.2011, leg. Assing (cAss). Antalya: 1 ex., SarÕcÕnar Da÷Õ, near Beldibi, 300 m, 6.-18.VI.1994, leg. Pütz (cAss). Ni ÷ d e: 1 ex., Çamardi env., Orhaniye, Ala Da÷lar, 1800-2200 m, 19.V.2005, leg. Vávra (cVav). Karaman: 1 ex. [det. Feldmann], N Ermenek, YÕldÕbel Geçidi, 36°49'N, 32°56'E, 1920 m, 24.V.2007, leg. Marggi & Huber (cFel). Israel: 1 ex. [det. Feldmann], S Hadera, Breikhat Ja'ar, 32°25'N, 34°54'E, 10 m, wet meadows, [date not specified] leg. Assmann (cFel). Ukraine: 1 ex., Odessa, Majaki, Dnestr river, at light, 18.-19.VIII.2005, leg Cibuƺskis (cSha) ; 4 exs., Odessa, Kuyalnik Liman, under stones, 2.X.2002, leg. Gontarenko (cAss). Georgia: 1 ex., Gyandzha (=Kirovabad) ["Elisabethpol"], Babadjanides (SMF). Azerbaijan: 7 exs., Geok Tapa [39°11'N, 48°41'E] (MHNG, cAss) ; 3 exs., Tejen ["Tedschen", 37°23'N, 60°30'E], V.1903, leg. Hauser (NHMW, cAss). Turkmenistan: 2 exs., Ashkhabad (SMF) ; 2 exs., W Ashkhabad, Sulyukly ["Transcaspia, Neu-Saratow", 38°02N, 57°25E] (MNHUB, cAss). Kazakhstan: 1 ex., Charyn valley, W Chundza, 800 m, 10.-13.VI.1993, leg. Schawaller (cAss). Locality not specified or illegible: 1 ex., " Transcaspien " (SMTD) ; 4 exs. (SDEI, MNHUB, SMTD) .

Comment: The vast distribution of S. anale ranges from southwestern France across the Mediterranean region to Iran and Middle Asia (Turkmenistan, Kazakhstan). In North Africa it has been reported only from Algeria (SMETANA 2004). The above specimens from Albania, Israel, Azerbaijan, Georgia, and Kazakhstan represent new country records; S. anale is not listed for these countries in SMETANA (2004). The species is usually collected from under stones in moist meadows, occasionally in large numbers particularly during spring inundations, often together with Leptobium gracile (GRAVENHORST 1802) and species of Luzea BLACKWELDER 1952 and/or Achenium LEACH 1819 . On one occasion, as many as 277 specimens were found in a partly inundated fallow near a road margin in Turkey (Fig. 1). For illustrations of the male sexual characters see COIFFAIT (1982).