Harpilius lutescens Dana, 1852

Harpilius lutescens Dana, 1852: 25

Harpilius consobrinus Balss, 1915: 27 (partim). — Bruce, 1992a: 72 –74, fig. 21 A–G.

Harpilius depressus — Tattersall, 1921: 389 –390, pl. 28 fig.7.

Harpilius ? lutescens — Kemp, 1922: 235 –237, figs 72–73.

Periclimenes (Ancylocaris) amamiensis Kubo, 1940a: 44 –46, figs. 11–12.

Periclimenes (Harpilius) lutescens — Holthuis, 1952: 88 –91, fig. 35.

Periclimenes lutescens — Bruce, 1972: 411, fig. 1A; 1992: 72–73, fig. 21a–g. — Chace and Bruce, 1993: 117 –118. — Li, 2000: 209 –21 1, fig. 271.

Type material. No longer extant.

Type locality. Tongatapu Island, Tongan Islands.

Hosts. Generally associated with corals of the genus Acropora [ Scleractinia]. Also reported from Pocillopora damicornis (De Grave, 2000) .

Habitat. Shallow water coral reefs.

Bathymetric range. 2–6 m (Bruce, 1976); 3 m (Bruce, 1981a).

Distribution. Also known from Saudi Arabia, Kenya, Tanganyika, Comoro Islands, Réunion, Indonesia, Thailand, Vietnam, Philippines, Western Australia, and Queensland.

Remarks. The discrepancy between the line illustrations and the colour figure provided by Dana (1855, pl. 37 fig.4) is discussed by Chace and Bruce (1993). Dana does not indicate how many specimens were available to him. It is clear that he was dealing with more than one species. The colour pattern illustrated corresponds sufficiently closely to that of Philarius imperialis (Kubo, 1940b) for that species to be considered as also occurring in the Tongan Islands. This species is also an associate of Acropora species and may be found in association with H. lutescens . Its colouration is illustrated by Bruce (1977a, p. 73; 1986, p. 162 fig. 2) and Minemizu (2000, p. 73). Harpilius lutescens has been illustrated in colour by Bruce (1975, p.26; 1977a, p. 73) and Minemizu (2000, p. 46). Tattersall (1921, as H. depressus) also illustrated the ambulatory dactyl and clearly shows that his specimen was not Harpiliopsis depresssa (Stimpson) . He was also the first author to comment on the presence of a median process on the fourth thoracic sternite in a pontoniine shrimp.