Genus Mesoptilus Lameere, 1917

Type species: Mesoptilus dolloi Lameere, 1917; Stephanian, Carboniferous (= Gzhelian, Upper Pennsylvanian, Commentry Basin, Allier department, France.

Species includes: The type species and Mesoptilus carpenteri sp. nov.

Emended diagnosis.— In forewing, a dense net of irregular cells in area between C and ScP; a broad area between CuA and CuP with a dense net of small irregular cells; only seven-eight main branches of RP; part of CuA basal of brace m-cua long.

Remarks.— Mesoptilus, already attributed to the “ Strephocladidae ” by Béthoux and Wieland (2009) has all the characters supporting an attribution to the Anthracoptilidae, plus the presence of a posterior branch of RA near its apex, as in Strephocladus permianus .

Stratigraphic and geographic range.—Stephanian, Carboniferous (= Gzhelian, Upper Pennsylvanian), France.