Anorus piceus LeConte, 1859

(Figs. 5–7)

Anorus piceus LeConte 1859a: 87 . Type locality: San Diego, California.

Anorus piceous [lapsus calami]: Blaisdell 1934: 320.

Diagnosis. As in A. parvicollis, males of this species (Fig. 5) are readily separated from Dascillus by the prosternal process forming a short denticle near the anterior coxal margin, resulting in the procoxal cavities being contiguous externally (prosternal process completely separating procoxae in Dascillus). It may be separated from A. parvicollis by the dense, recumbent setae present throughout the dorsal surface, including the scutellar shield, and lack of long erect setae, though many specimens have a pelage of suberect setae on the elytra (A. parvicollis bearing sparse, long, mostly erect setae). The pronotum is distinctly narrowest across the anterior angles, then the pronotum gradually widens posteriorly for two-thirds of the length of pronotum, then narrows slightly to the posterior angles (Fig. 5A). The elytron is more similar to that of Dascillus than to that of A. parvicollis, both in disposition of setae and the presence of a sulcus along the lateral margin extending from behind the humerus nearly to the apical angle. The mandibles are more elongate in this species than A. parvicollis, the incisor edge is longitudinally straight anterior to the labrum, and there are two retinacula (proximal one longer and more acute) before the apex. Females are easily recognized by the disposition of the elytra, which extend about a third to a half of the distance to the tip of the abdomen (Fig. 6; see also Blaisdell 1934: fig. 3).

Redescription of Male. Body parallel-sided, moderately slender, 8–14 mm (Fig. 5A). Body setae of two types, most fairly short and recumbent; elytra, especially laterally, and appendages also bearing scattered moderately long, bristling setae. Antennomeres 3–10 weakly serrate, at least 2 times longer than wide. Left mandible bearing two strong retinacula, lacking basal angulation. Right mandible with two retinacula, basal retinaculum very strong and finger-like, more apical retinaculum blunt, appearing as a lobe on the incisor edge near the base of the apical tooth and often nearly absent. Epicranial suture faintly traceable. Lateral sides of pronotum weakly explanate, lateral carina always distinct to anterior margin, posterior margin weakly but distinctly bisinuate; pronotum widest in posterior third, anterior margin always narrower than posterior margin. Hypomeron with distinct but short postcoxal process. Prosternal process acute apically, lateral margins converging near middle of procoxae, continued as thin vertical lamina often concealed by procoxae (as in Fig. 6B); mesoventrite with abrupt mesal elevation projecting anteriorly, bearing small procoxal rests. Elytra attaining the apex of abdomen, with weakly punctate striate. Hind wing exceeding apex of abdomen; radial cell completely closed proximally; wedge cell medium-sized, separated from posterior edge by distance about half length of cell; apical field about one-third length of wing. All tibiae lacking stout spinules along outer margin. All legs with tarsomeres 1–4 bearing fleshy bilobed pads beneath, tarsomere 1 with relatively small lobes. Terminalia with tergite IX (Fig. 5G) with very broad and shallow V-shaped emargination along posterior margin; tergite X (Fig. 5G) evenly broadly rounded apically; sternite IX (Fig. 5F) rounded apically; phallobase of aedeagus (Figs. 5B–D) weakly emarginate to rounded basally, slightly broader than long; parameres narrow, much longer than phallobase, gradually narrowing from base to apex, lateral margin slightly concave, recurved apically, bearing very few scattered, fine setae in apical half; dorsal lobe of penis comparatively narrow, measured from anterior margin of basal strut to posterior apex approximately as long as paramere, variably narrowed apically, apex evenly to narrowly rounded and never appearing cleft; ventral lobe of penis nearly as long as dorsal lobe, curved dorsally around middle, then curved ventrally near apex, apex narrowly rounded.

Redescription of Female. Resembling male in most characters except: Body stouter (Fig. 6), wider in apical half. Elytra shorter than abdomen, about 1.5–2× as long as pronotum, lacking striae. Hind wing less than half length of abdomen. Terminalia (Fig. 7) with tergite IX not observed; sternite VIII (spiculum ventrale) extremely weakly lobed medially; proctiger narrowly triangular; paraprocts short, with ventral baculi converging apically; gonocoxites slightly shorter than paraprocts, each coxite with proximal plate more heavily sclerotized; gonostyli apparently absent.

Distribution. Primarily along the drier slopes and valleys of the Coast Ranges in southern California but extending at least to Owens Valley (Inyo Co.) to the northwest and south into Baja California (Fig. 8).

Note on Variation and Type Material. LeConte (1859a) did not mention how many specimens his original description was based on. Four male specimens are deposited in the LeConte collection (MCZ), three of which bear a gold circle (= “ California ”); the latter should probably be considered syntypes.

The examined specimens of A. piceus demonstrate less individual and inter-populational variation than is seen in A. parvicollis. Specimens of the former from California and Baja California tend to be similar in terms of coloration and sclerotization. Specimens from the northeastern extent of its range in Nevada tend to be slightly smaller and lighter in color, and the female elytra are proportionately shorter than those of the single female known from California.

Specimens Examined (671). UNITED STATES: CALIFORNIA: Inyo Co.: 7 miles west of Westgard Pass, 25 June 1953, H. Washburn (1♂, LACM); Alabama Hills, 36.60083°, 118.10615°, 1,350 m, 1 June 2016, UV/MV light, M. and L. Gimmel, A. Mayor, M. A. Johnston, C. Wirth, ML83 (1♂, SBMNH); Antelope Spring, 8 mi SW of Deep Springs, 29 June 1961, J. A. Litsinger (1♂, MAIC); same except 9 July 1962 (1♂, MAIC); Argus Mountains, 8 June 1938, J. A. Comstock (6♂♂, LACM); Baker Creek Campground, Big Pine, 37° 10.180 N, 118° 18.550 W, 8 June 2011, M. Kippenhan (1♂, MAIC); Big Pine Canyon, 8,5000, 8 August 1973, G. R. Ballmer (2♂♂, UCRC); Independence, 6 June 1918, L. L. Muchmore (2♂♂, LACM); Independence, 1 June 1972 (4♂♂, UCRC); Junction Ranch, 28 June 1969, L. Lester (2♂♂, LACM); Wyman Canyon, White Mountains, 9 July 1967, S. and S. Fromer (3♂♂, UCRC); 5 mi W of Big Pine, Baker Creek, 37.15424°N, 118.34295°W, 4,939 ft, 17 May 2004, UV light and trap, Thomas E. Dimock, 1121 (4♂♂, SBMNH); Kern Co.: Democrat Hot Springs, 2,2000 elevation, 16 June 1969, D. Levin (1♂, LACM); Keene, 3,000 ft, 7 July 1975, A. Fuller (1♂, SBMNH); Rancheria Creek in Piute Mountains, 4,3500 elevation, T29S R33E S23, 1 June 1973, J. P. and K. E. Donahue (2♂♂, LACM); Los Angeles Co.: 1 mile south of Valyermo, 19 May 1964, M. J. Wargo (3♂♂, UCRC); 2.5 miles SSW of Valyermo, 20 June 1959, N. McFarland (5♂♂, LACM); 5 miles north of Beverly Hills in Santa Monica Mountains, 1,1000 elevation, 13 July 1955, N. McFarland (3♂♂, LACM); Albright (2♂♂, LACM); Altadena, 10 May 1959, J. R. Northern (1♂, LACM); Angeles National Forest, San Dimas Experimental Forest, Bell Canyon, 34.1826°N, 117.7938°W, 13 June 2007, M. Caterino and S. Chatzimanolis (5♂♂, SBMNH); Angeles National Forest, Ruby Canyon, 34.6060°N, 118.5523°W, 22 June 2007, at light, M. Caterino and S. Chatzimanolis (4♂♂, SBMNH); Angeles National Forest, San Dimas Experimental Forest, Big Dalton Canyon, 34.1793°N, 117.7985°W, 27 July 2007, M. Caterino (1♂, SBMNH); Angeles National Forest, Shake Canyon, 34.7009°N, 118.5250°W, 2 July 2007, at light, M. Caterino (1♂, SBMNH); Big Pine Ranger Station, 5 July 1975, A. V. Evans (16♂♂, LACM); Big Pines, 13 July 1971 (6♂♂, LACM); Big Pines Recreation Area, 7 July 1983, S. Ziff (3♂♂, LACM); Big Pines Recreation Area, 20 July 1983, S. Ziff (2♂♂, LACM); Bouquet Canyon, 23 June 1937, N. Westerlund (3♂♂, LACM); Bouquet Canyon, 26 June 1937, J. A. Comstock (9♂♂, LACM); Bouquet Canyon, 7 July 1937, N. Westerlund and L. M. Martin (16♂♂, LACM); Burbank, 21 May 1930, C. H. Hicks (1♂, CASC); Burbank, 23 May 1930, C. H. Hicks and F. C. Hadden (3♂♂, LACM); Burbank, 30 June 1958 (2♂♂, LACM); Chatsworth, 26 June 1920, L. L. Muchmore (2♂♂, LACM); Claremont, 19 May 1962, T. Reimo (2♂♂, UCRC); Devil’ s Punch Bowl, 21 June 1964, D. G. Marqua and P. H. Sullivan (1♂, LACM); Devil’ s Punchbowl, 11 June 1975 (15♂♂, LACM); Eaton Canyon Park, 5 June 1964, R. H. Crandall (2♂♂, LACM); Glendale, 3 July 1965, E. I. Schlinger (1♂, UCRC); Gorman, 22 July 1938, J. von Bloeker (6♂♂, LACM); Juniper Hills, 8 June 1975, A. V. Evans (23♂♂, LACM); Juniper Hills, 4,3000 elevation, 14 July 1979, C. L. Bellamy (2♂♂, LACM); La Canada, 8 July 1950, D. C. Blodget (3♂♂, LACM); La Canada, 21 June 1969 (1♂, UCRC); La Crescenta, 19 June 1955, J. Roberds (6♂♂, LACM); La Verne, 24 June 1961, R. S. Erdman (4♂♂, LACM); Lake Hughes, 13 July 1938 (1♂, LACM); Littlerock, [day/month blank] 1945 (1♂, LACM); Littlerock, [day blank] May 1947, G. P. Mackenzie (1♂, LACM); Mint Canyon, 21 June 1949, H. E. Cott (1♂, SBMNH); Mount Wilson (2♂♂, LACM); Pomona, 6 April 1931, W. D. Clancy (1♂, UCRC); Santa Monica, 22 July 1943, G. P. Mackenzie (1♂, UCRC); Sierra Madre, 11 July 1944 (1♂, LACM); Sunland, 12 May 1940 (1♂, LACM); Tanbark Flats, 4 July 1950, D. C. Blodget (1♂, LACM); Tujunga, 19 June, 1955, J. Roberts (1♂, LACM); Tujunga, 14 May 1969, J. C. von Bloeker (30♂♂, LACM); Tujunga Canyon, 15 May 1969, A. Strong (1♂, UCRC); Littlerock, 8 June 1941, G. P. Mackenzie (1♂, UCRC); Topanga Canyon, 6 June 1978 (1♂, LACM); Pasadena, 25 May 1894 (1♂, LACM); Pasadena, [day blank] May [year blank] (1♂, LACM); San Marino, 5 July 1942, G. P. Mackenzie (1♂, UCRC); Valyermo, 1 July 1967, S. Ziff (2♂♂, LACM); West Los Angeles, 17 March 1967, B. S. Cheary (1♂, UCRC); [county record only], [day blank] June 1938, J. Schwartz (1♂, LACM); Orange Co.: 4 miles east of Olive, 1 July 1975, J. Wilcox (1♂, UCRC); Orange Park, 11 May 1978, B. P. Harris (1♂, LACM); Riverside Co.: 4 miles south of Beaumont, at Potrero Creek, 1 July 1990, G. R. Ballmer and D. C. Hawks (1♂, UCRC); 6 kilometers north of Desert Center, 18 March 1986, W. E. Steiner and J. E. Lowry (2♂♂, USNM); Beaumont, 10 June 1942, E. Van Dyke (1♂, CASC); Bee Canyon near Strawberry Canyon, 25 May 1969 (♂, LACM); Box Spring Mountains, 30 June 1964, G. E. Wallace (1♂, UCRC); same except C. D. Howell (4♂♂, UCRC); Big Morongo, 22 April 1985, M. Clement (3♂♂, UCRC); Big Morongo Canyon, 2,0000 elevation, 2 May 1969, C. D. Howell (3♂♂, UCRC); Boggert Trail, T 5S R4E S14, 25 April 1998, J. D. Barry (5♂♂, UCRC); Bradshaw Trail Road 4 miles west of Graham Pass road, 33° 25.60 N 115° 14.40 W, 11 April 1994, R. R. Snelling (5♂♂, LACM); Citrus Experimental Station, 13 May 1964, P. A. Rauch (1♂, UCRC); Corona, 5 June 1955, R. H. Alvarado (1♂, UCRC); Deep Canyon, Sec. 6 T 7s, R 6E, 16 May 1975, L. E. Clark (9♂♂, UCRC); Dripping Springs, 33° 270 N 116° 580 W, 13 May 1999 (3♂♂, UCRC); Gavilan Hills, 15 May 1965 (11♂♂, UCRC); Indian Wells, 24 April 1953, G. Yamamoto (1♂, LACM); Lake Elsinore, 28 May 1981, G. Pratt (2♂♂, UCRC); Lost Palms Canyon in Joshua Tree National Monument, 26 May 1963, E. Sleeper (1♂, CASC); Morongo Valley 1 mile north of Indian Avenue, 3 May 1969, B. Hunt (2♂♂, UCRC); P. L. Boyd Desert Research Center, 3.5 miles south of Palm Desert, 3 May 1974, D. L. Dickson (1♂, UCRC); Pinyon Flats, 6 June 1970, Cazier, Francke and Welch (12♂♂, ASUHIC); Pinyon Flats in San Jacinto Mountains, 27 May 1939, J. G. Shanafelt (2♂♂, LACM); Pinyon Flats in San Jacinto Mountains, 27 June 1952, B. Tinglof (2♂♂, LACM); Pinyon Flats, 4,0000 elevation, 26 May 1980, C. L. Bellamy (1♂, LACM); Riverside, 2 May 1961, L. D. Anderson (1♂, UCRC); Riverside, 5 May 1961, O. L. Brawner (2♂♂, UCRC); Riverside, 2 May 1961, C. S. Papp (5♂♂, LACM); Riverside, 1 May 1962, B. Ewing (4♂♂, UCRC); Riverside, 7 May 1962, E. I. Schlinger (3♂♂, UCRC); Riverside, 26 June 1962 (1♂, UCRC); Riverside, 14 May 1964, M. E. Irwin (1♂, UCRC); Riverside, 29 May 1964, M. E. Irwin (1♂, UCRC); Riverside, 13 May 1972, A. Tabet (8♂♂, UCRC); Riverside, 16 May 1972, A. Tabet (3♂♂, UCRC); Riverside, 23 May 1972, A. Tabet (2♂♂, UCRC); Riverside, 2 June 1972, A. Tabet (2♂♂, UCRC); San Bernardino National Forest, San Jacinto River, 33.7244°N, 116.8061°W, 24 May 2005, at light, M. Caterino (5♂♂, SBMNH); Split Rock, Joshua Tree National Monument, 24 May 1966, J. Wilcox (3♂♂, UCRC); Tahquitz Canyon, 8 June 1957, Menke and Stange (2♂♂, LACM); Temecula Canyon, Santa Margarita River, 7500 elevation, 18 May 1968, L. O. Tejada (1♂, UCRC); Tenaja Campground, 14 May 1972, A. Tabet (3♂♂, UCRC); Tenajas Ranger Station, 29 July 1967, J. A. Honey (3♂♂, LACM); Hurkey Creek Campground, 2 May 1972 (1♂, USNM); Whitewater Canyon, 16 May 1969, B. Duff (1♂, USNM); Whitewater Canyon, 33° 6202200 N 116° 4104200 W, J. D. Barry (1♂, UCRC); Whitewater Canyon, T 2S R3E S 26, 650 m elevation, 6 May 1997, H. W. Park (3♂♂, UCRC); [county record only] 13 May 1966, M. W. Stone (3♂♂, UCRC); 24 May 1966, M. W. Stone (3♂♂, UCRC); Whitewater Canyon, 33.9692°N, 116.6521°W, 15 May 2006, M. Caterino and S. Chatzimanolis (6♂♂, SBMNH); San Bernardino Co.: 44 km S of Needles, 34.4388°N, 114.6517°W, 28 April–4 May 2007, Malaise trap in dry wash with flowering Cercidium, M. E. Irwin (3♂♂, SBMNH); Alta Loma, 11 June 1955, R. H. Alvarado (2♂♂, UCRC); Apple Valley, 8 July 1959, N. McFarland (1♂, LACM); Baldy Mesa, 13 June 1937, J. A. Comstock (2♂♂, LACM); Cajon Pass, 24 June 1937 (1♂, LACM); Cajon Pass, 24 June 1949, H. E. Cott (1♂, SBMNH); Clark Mountain, T 17N R13E SE quarter S22, 25 May 1985, J. P. and K. E. Donahue (3♂♂, LACM); Cucamonga Canyon, 22 May 1972, T. Stemwedel (1♂, LACM); Deep Creek Campground, 15 June 1937, Menke and Stange (4♂♂, LACM); Fontana, 19 May 1959, R. J. Jeffrey (2♂♂, UCRC); same except 20 May 1959 (2♂♂, UCRC); Horse Spring Campground in San Bernardino Mountains, 5700’ elevation, 13 June 1986, J. P. and K. E. S. Donahue (6♂♂, LACM); Joshua Tree National Park, 49 Palms Oasis Trail, 34.1109°N, 116.1072°W, 27–28 April 2012, UV light, M. S. Caterino, A. K. Tishechkin and M. P. Richmond (4♂♂, SBMNH); Joshua Tree National Park, 49 Palms Oasis Trailhead, 34.1193°N, 116.1118°W, 27–28 April 2012, D. Yanega (1♂, SBMNH); Joshua Tree National Park, 49 Palms Oasis, 34.1058°N, 116.1051°W, 27–28 April 2012 (1♂, SBMNH); Joshua Tree National Park, Smithwater Canyon, 34.0213°N, 116.2961°W, 25–27 April 2013, night collecting, M. S. Caterino and E. Shafer (1♂, SBMNH); Lower Covington Flat in Joshua Tree National Monument, 28 May 1960, E. Sleeper (1♂, CASC); Lytle Creek, 8 June 1928, E. Van Dyke (1♂, CASC); Mill Creek, 957 m elevation, 31 May 1994, M. Cooperband (1♂, UCRC); Oak Glen, 2 June 1970, M. J. Wargo (1♂, UCRC); Ontario, 1 June 1938 (1♂, UCRC); Pioneertown, UC Burns Reserve, 1350 m elevation, 2 August 1994, J. Freilich (1♂, UCRC); Pioneertown, UC Burns Reserve, 1350 m elevation, 28 May 1995, J. Freilich (1♂, UCRC); Pioneertown, UC Burns Reserve, 1350 m elevation, 1 July 1995, J. Freilich (2♂♂, UCRC); Providence Mountains, 7 May 1939, J. A. Comstock (2♂♂, LACM); San Bernardino National Forest, Gobbler’ s Knob, 34.3116°N, 117.5835°W, 3 July 2005, M. Caterino (3♂♂, SBMNH); Seven Oaks, 8 July 1950 (1♂, LACM); Twentynine Palms, 26 April 1949 (2♂♂, LACM); UC Burns Reserve, 34.1403°N, 116.4542°W, 22 May 2005, at light, M. Caterino (1♂, SBMNH); Victorville, 20 May 1935, C. Dammers (2♂♂, LACM); Wildwood Canyon, 5 miles east of Calimesa, 28 July 1965, E. I. Schlinger (1♂, UCRC); Wrightwood, 19 June 1966, R. H. Crandall (6♂♂, LACM); Wrightwood, 20 July 1983, S. Ziff (38♂♂, LACM); Wrightwood, 2 June 1984, S. Ziff (1♂, LACM); San Diego Co.: Alpine, 20 April 1960, S. C. Williams (2♂♂, ASUHIC); Borrego, 5 April 1940 (3♂♂, LACM); Borrego, 27 April 1955, Timberlake (1♂, UCRC); Boulder Oaks, 30 June 1957, D. Peterson (2♂♂, UCRC); Boulevard near I-8 exit, 3 June 1995 (1♂, MAJC); Culp Canyon, 2 April 1962, J. C. Hall (1♂, UCRC); Encinitas, 26 May 1932, H. L. McKenzie (3♂♂, UCRC); Indian Flats Campground, 3,8000 elevation, 28 May 1968, J. A. Honey (9♂♂, LACM); Jacumba, [day/month blank] August 1955, D. Cox (1♂, ASUHIC); La Puerta Valley, 23 April 1938 (4♂♂, LACM); La Puerta Valley, 20 May 1939, L. M. Martin (3♂♂, LACM); Lawson Valley, 12 June 1939, J. von Bloeker (4♂♂, LACM); Oak Grove, 28 June 1966, J. R. Dixon (2♂♂, LACM); Palm Canyon, Borrego Valley, 28 May 1936, Timberlake (15♂♂, UCRC); Palm Canyon, Borrego Desert, 2 April 1946, A. L. Melander (1♂, UAIC); Palm Canyon, Borrego Valley, 26 April 1953 (2♂♂, LACM); Palomar Mountain, 22 June 1940, G. P. Mackenzie (1♂, UCRC); Point Loma, 17 June 1952, R. V. D. Rosch (1♂, UCRC); Scissors Crossing, Valle de San Felipe, 2,240 – 2,8000 elevation, 4 May 1969, E. I. Schlinger (2♂♂, UCRC); Sentenac Canyon, 26 April 1953, G. Yamamoto (1♂, LACM); Sentanac Canyon, 1,800 – 1,9000 elevation, 4 May 1968, J. DeVol (1♂, UCRC); Torrey Pines, 6 July 2009, N. Bloomfield (1♂, USNM); [county record only], F. E. Blaisdell (1♀, CASC); [county record only] 10 June 1966, M. W. Stone (1♂, UCRC); 27 June 1967, M. W. Stone (2♂♂, UCRC); San Luis Obispo Co.: 7 mi E of Pozo, 4 July 1967, blacklight, C. W. Baker and D. H. Wilson (1♂, SBMNH); Carrizo Plain National Monument, 35.0440°N, 119.6009°W, 24 May 2003, mercury vapor light, M. Caterino and P. Jump (13♂♂, SBMNH); Carrizo Plain National Monument, Selby Campground, 35.1273°N, 119.8430°W, 5 June 2004, at light, M. Caterino and P. Jump (3♂♂, SBMNH); Los Padres National Forest, 27 June 1987, L. Hunsaker (1♂, SBMNH); Mountains west of La Panza, 18 May 1928, J. O. Martin (4♂♂, CASC); Paso Robles, 24 May 1928, L. S. Slevin (1♂, CASC); Paso Robles, 12 June 1971, C. E. Langstrom (1♂, LACM); Santa Margarita, 12 June 1971, C. E. Lengstrom (4♂♂, LACM); Santa Barbara Co.: 0.75 mi S of Ballinger Wash, 34.876°N, 119.482°W, 2,840 ft, 13 June 2007, UV light trap, P. M. Jump (3♂♂, SBMNH); same except 20 June 2007 (5♂♂, SBMNH); same except 7 June 2007 (2♂♂, SBMNH); Ballinger Canyon, check dam, 34.87665°, 119.48225°, 860 m, 6 May 2017, UV light trap, Peter Jump (3♂♂, SBMNH); state record only: May 1908, R. Woglum (1♂, UCRC). NEVADA: Clark Co.: Charleston Peak, 4 June 1946 (3♂♂, CASC); Nye Co.: Mercury Nevada Test Site, 18 May 1960 (1♂, MAIC; 6♂♂, USNM); same except 19 May 1960 (3♂♂, USNM); same except 19 May 1961 (3♂♂, USNM); same except 25 May 1961 (1♂, MAIC; 3♂♂, USNM); same except 7 June 1961 (1♂, USNM); same except 16 June 1964 (1♀, USNM); same except 20 June 1964 (1♂, 2♀♀, MAIC; 1♂, 3♀♀, USNM); same except 12 June 1965 (1♂, USNM). MEXICO: BAJA CALIFORNIA [NORTE]: 4 miles south of La Rumorosa, 4,4000 elevation, 7 July 1967, M. E. Irwin (5♂♂, UCRC); 12 miles east of El Rosario, 10 June 1979 (4♂♂, UCRC); Arroyo del Rosario 3 miles above Rosario, 26 April 1963, P. H. Arnaud (5♂♂, CASC); La Zapopita, Valle de Trinidad, 7 April 1961, F. S. Truxal (7♂♂, LACM); La Zapopita, Valle de Trinidad, 16 April 1961, F. S. Truxal (9♂♂, LACM); Parque Nacional Constitución de 1857, 12 km S of Laguna Hanson, Sierra Juárez, 31.96°N, 115.99°W 1,700 m, 15 June 2019, A. Shamaev (3♂♂, MAIC); San Telmo Canyon in San Pedro Martir, 30 May 1981, Werner, Olson, Hetz, Thomas, Burn, Frank and MacLachlan (4♂♂, UAIC); Santo Tomas, 8 August 1973, S. C. Williams and K. B. Blair (2♂♂, CASC); Tijuana, 18 June 1964 (1♂, UCRC). Doubtful localities, not mapped: CALIFORNIA: Yolo Co.: Davis, 10 May 1960, J. L. Bath (2♂♂, UCRC). BAJA CALIFORNIA SUR: Santa Inez, 26 April 1979, D. B. Weissman (1♂, CASC) [probably in Baja Norte or mislabeled]. WYOMING: Converse Co.: Douglas, 27 July 1964, H. B. Leech (1♂, CASC) [almost certainly mislabeled].