Gonatocerus (Cosmocomoidea) novifasciatus Girault, 1911

(Figs 417–423)

Gonatocerus novifasciatus Girault 1911: 266 –267. Holotype female [USNM] (not examined). Type locality: an unspecified locality in Tennessee, USA.

Lymaenon novifasciatus (Girault): Burks 1958: 63 (catalog); Turner & Pollard 1959: 26 (host association); Peck 1963: 24 (catalog).

Gonatocerus novifasciatus Girault: Huber 1988: 63 –65 (taxonomic history, redescription, diagnosis, distribution, host associations); Yoshimoto 1990: 40 (list, ater species group); De Santis & Fidalgo 1994: 125 (catalog); Triapitsyn et al. 1998: 241, 243 (host association in California); Triapitsyn et al. 2006: 62–63 (molecular data); Triapitsyn 2006a: 27 (illustration), 28–30 (diagnosis, distribution, host associations); de León et al. 2006a: 42 (molecular data); de León et al. 2006d: 54 (molecular data); de León et al. 2006e: 57–58 (molecular data); Bernal et al. 2007: 67 –68 (record from Sonora, Mexico); de León & Morgan 2007: 83 (molecular data); de León et al. 2008: 99, 105 (molecular data); Triapitsyn & Bernal 2009: 7 (host associations).

Material examined. COLOMBIA. MAGDALENA, Parque Nacional Natural Sierra Nevada de Santa Marta, La Estación, 10°48’N 73°39’W, 2220 m, 15–31.vii.2000, J. Cantillo [2 ♀, UCRC] . GUATEMALA. SACATEPÉQUEZ, Sumpango, Durwest Farm, 14°40.292’N 90°43.195’W, 5985’, 1–13.xii.2007, M.S. Hoddle [2 ♀, UCRC] . MEXICO. VERACRUZ, Orizaba, 28.vi.1966, J.S. Buskets, N.R. & R.C. Gardner [1 ♀, UCDC] .

Extralimital records. USA. ARIZONA, Graham Co., Coronado National Forest, Snow Flat, 32°39’15’’N 109°51’49’’W, 2690 m, 19–20.viii.2001, J.B. Woolley [1 ♀, UCRC] . CALIFORNIA: Los Angeles Co.: Altadena, 1.iv.1963, R.H. Crandall [1 ♀, LACM]. Santa Monica Mountains, Big Rock Road, 34°02’N 118°37’W, 12-18.vi.1997, M.E. Irwin [3 ♀, 4 ♂, UCDC]. Monterey Co., Monterey State Beach North (near Seaside), 36°37’N 121°51’W, 8.vi.2005, S.L. Heydon [1 ♀, UCDC] . MISSISSIPPI, Washington Co., Delta Experimental Forest (near Stoneville), 33°27’N 90°55’W, 25.v-8.vi.1998, N.M. Schiff [1 ♀, UCDC] . NEW YORK, Onondaga Co., Jamesville, Henneberry Rd., 42°55’00’’N 76°00’23’’W, 17–27.vii.2001, M. Wuenschel [1 ♀, UCRC] . PENNSYLVANIA, Montgomery Co., Lansdale, 1108 Amber Lane, 40°14’12’’N 75°18’47’’W, 20–27.ix.2001, R. Kaufhold [1 ♂, UCRC].

Redescription. See Huber (1988).

Diagnosis. Gonatocerus (Cosmocomoidea) novifasciatus is a member of the ater subgroup of the ater species group. Its main distinguishing features are as follows. Body dark brown. Female antenna (Fig. 417) with mps usually present on F3–F8 (F4 rarely lacking mps), F1 and F2 usually without mps (F2 rarely with 1 mps). Male antenna (Fig. 420) with F1 about as long as wide. Propodeum (Figs 418, 421) with submedian carinae uniformly thickened, united anteriorly at the dorsellum to form a closed median area on the propodeum. Forewing (Figs 419, 423) hyaline or with faint, uniform, brown tinge, and with usually distinct, occasionally indistinct, infumate spot just beyond the stigmal vein (Neotropical specimens tend to have a somewhat darker spot than Nearctic ones); discal setae absent behind venation except for a few setae behind apex of the stigmal vein. Male genitalia as in Fig. 422.

Distribution. NEOTROPICAL: Colombia *, Guatemala *, Honduras, Mexico (Triapitsyn 2006a), and Venezuela (Huber 1988). NEARCTIC: Canada (Huber 1988), Mexico (Triapitsyn 2006a), and USA. OCEA- NIA: Hawaiian Islands (USA: Hawaii) (Triapitsyn 2006a).

Hosts. Graphocephala coccinea (Förster), G. versuta (Say), Homalodisca liturata Ball, H. vitripennis (Germar) (Huber 1988; Triapitsyn 2006a), and also very likely (although not yet documented) Oncometopia orbona (Fabricius) (Triapitsyn 2006a) ( Cicadellidae).