3. Labiobaetis glaucus (Agnew, 1961)

Baetis glaucus Agnew 1961 ( Baetis glaucus)

Labiobaetis masai Lugo-Ortiz and McCafferty 1997: figs 27-38, 39-50 ( Labiobaetis masai, L. nadineae; both formal synonyms, Lugo-Ortiz et al. 2000)

Pseudocloeon glaucum Lugo-Ortiz et al. 2000 ( Pseudocloeon glaucum)

Labiobaetis glaucus Gattolliat et al. 2018: figs 34-44, 47

Differential diagnosis.

Nymph. Following combination of characters: A) colouration: abdomen dorsally brown, with pattern as Gattolliat et al. 2018: fig. 47; B) scape without distolateral process; C) labial palp segment II with broad thumb-like protuberance; D) maxillary palp segment II with excavation at inner distolateral margin; E) fore femur rather broad, length ca. 3 × maximum width; dorsal margin with 13-18 curved, spine-like setae and basally some additional setae near margin; femoral patch well developed; F) fore tibia dorsally with a row of scarce, tiny, stout setae (Gattolliat et al. 2018: fig. 40); G) hind protoptera well developed; H) seven pairs of gills; I) paraproct with 5-10 stout, marginal spines.

Examined material.

Ethiopia • 6 nymphs; Middle Borkana River; 10°38'09"N, 39°55'53"E; 17.03.2019; 1413 m; leg. W. Graf; 1 on slide; GenBank MW307230; GBIFCH00763728; 5 in alcohol; GBIFCH00515556 • 4 nymphs; Lafessa; 08°23'16"N, 38°54'31"E; 1600 m; 08.11.2017; leg. W. Graf; Lf1; in alcohol; GBIFCH00515557 • 1 nymph; Dubti; 11°41'50"N, 41°07'23"E; 2017; 374 m; leg. W. Graf; S14; in alcohol; GBIFCH00515564 • 1 nymph; Sulula; 08°39'57"N, 38°37'59"E; 1916 m; 07.11.2017; leg. W. Graf; Su1; in alcohol; GBIFCH00515563 • 2 nymphs; Worer; 09°20'6.98"; 40°10'19.50"; 740 m; 29.01.2018; leg. W. Graf; Wr1; 1 on slide; GBIFCH00592437; 1 in alcohol; GBIFCH00515565; all material in MZL .

Biological aspects.

The specimens were collected at altitudes from 370 m to 1920 m. Further characteristics of sampling sites are given in Englmaier et al. (2020). Harrison and Hynes (1988) reported the species from 750 m to 1900 m in stony runs and torrents. In Saudi Arabia, the species occurs in small, very shallow streams with moderate current and a substrate mixed of sand, cobbles and rock (Gattolliat et al. 2018).

Distribution.

Ethiopia (Fig. 2a; Harrison and Hynes 1988), Saudi Arabia, Comoros (Gattolliat et al. 2018), South Africa, Lesotho, Namibia, Kenya (Lugo-Ortiz et al. 2000), Zimbabwe (Harrison and Hynes 1988) and potentially Iran (Tahmasebi et al. 2020).